THE CENTER FOR NORTH AMERICAN HERPETOLOGY


Provincial/State Species Lists and References

Canada: Alberta (20) British Columbia (41) Labrador (7) Manitoba (24) New Brunswick (24) Newfoundland (7) Northwest Territories (6) Nova Scotia (24) Nunavut (3) Ontario (53) Prince Edward Island (14) Quebec (38) Saskatchewan (22) Yukon (4)

USA:
Alabama (177) Alaska (10) Arizona (155) Arkansas (126) California (206) Colorado (74) Connecticut (50) Delaware (67) District of Columbia (59) Florida (192) Georgia (184) Hawaii (33) Idaho (40) Illinois (106) Indiana (97) Iowa (70) Kansas (99) Kentucky (114) Louisiana (150) Maine (39) Maryland (94) Massachusetts (53) Michigan (55) Minnesota (51) Mississippi (153) Missouri (117) Montana (34) Nebraska (62) Nevada (79) New Hampshire (42) New Jersey (79) New Mexico (136) New York (73) North Carolina (168) North Dakota (28) Ohio (83) Oklahoma (143) Oregon (72) Pennsylvania (81) Rhode Island (44) South Carolina (142) South Dakota (46) Tennessee (142) Texas (240) Utah (76) Vermont (40) Virginia (150) Washington (51) West Virginia (90) Wisconsin (55) Wyoming (40)



Established Extant Amphibians and Reptiles of Pennsylvania

Amphibia Gray, 1825 — Amphibians
Anura Fischer von Waldheim, 1813 — Frogs
Bufonidae Gray, 1825 — True Toads
Anaxyrus americanus (Holbrook, 1836) — American Toad
Anaxyrus fowleri (Hinckley, 1882) — Fowler's Toad
Hylidae Rafinesque, 1815 — Treefrogs and Allies
Acris crepitans Baird, 1854 — Eastern Cricket Frog
Dryophytes chrysoscelis (Cope, 1880) — Cope's Gray Treefrog
Dryophytes versicolor (LeConte, 1825) — Gray Treefrog
Pseudacris brachyphona (Cope, 1889) — Mountain Chorus Frog
Pseudacris crucifer (Wied-Neuwied, 1838) — Spring Peeper
Pseudacris feriarum (Baird, 1854) — Upland Chorus Frog
Pseudacris kalmi Harper, 1955 — New Jersey Chorus Frog
Pseudacris triseriata (Wied-Neuwied, 1838) — Western Chorus Frog
Ranidae Rafinesque, 1814 — True Frogs
Lithobates catesbeianus (Shaw, 1802) — North American Bullfrog
Lithobates clamitans (Latreille in Sonnini de Manoncourt and Latreille, 1801) — North American Green Frog
Lithobates kauffeldi (Feinberg, Newman, Watkins-Colwell, Schlesinger, Zarate, Curry, Shaffer, and Burger, 2014) — Atlantic Coast Leopard Frog
Lithobates palustris (LeConte, 1825) — Pickerel Frog
Lithobates pipiens (Schreber, 1782) — Northern Leopard Frog
Lithobates sphenocephalus (Cope, 1886) — Southern Leopard Frog
Lithobates sylvaticus (LeConte, 1825) — Wood Frog
Scaphiopodidae Cope, 1865 — North American Spadefoots
Scaphiopus holbrookii (Harlan, 1835) — Eastern Spadefoot
Caudata Scopoli, 1777 — Salamanders
Ambystomatidae Gray, 1850 — Mole Salamanders
Ambystoma jeffersonianum (Green, 1827) — Jefferson Salamander
Ambystoma laterale Hallowell, 1856 — Blue-spotted Salamander
Ambystoma maculatum (Shaw, 1802) — Spotted Salamander
Ambystoma opacum (Gravenhorst, 1807) — Marbled Salamander
Ambystoma tigrinum (Green, 1825) — Eastern Tiger Salamander
Cryptobranchidae Fitzinger, 1825 — Giant Salamanders
Cryptobranchus alleganiensis (Sonnini de Manoncourt and Latreille, 1801) — Hellbender
Plethodontidae Gray, 1850 — Lungless Salamanders
Aneides aeneus (Cope and Packard, 1881) — Green Salamander
Desmognathus fuscus (Green, 1818) — Northern Dusky Salamander
Desmognathus monticola Dunn, 1916 — Northern Seal Salamander
Desmognathus ochrophaeus Cope, 1859 — Allegheny Mountain Dusky Salamander
Eurycea bislineata (Green, 1818) — Northern Two-lined Salamander
Eurycea longicauda (Green, 1818) — Long-tailed Salamander
Gyrinophilus porphyriticus (Green, 1827) — Spring Salamander
Hemidactylium scutatum (Temminck, 1838) — Four-toed Salamander
Plethodon cinereus (Green, 1818) — Eastern Red-backed Salamander
Plethodon electromorphus Highton, 1999 — Northern Ravine Salamander
Plethodon glutinosus (Green, 1818) — Northern Slimy Salamander
Plethodon hoffmani Highton, 1972 — Valley and Ridge Salamander
Plethodon wehrlei Fowler and Dunn, 1917 — Wehrle's Salamander
Pseudotriton montanus Baird, 1850 — Mud Salamander
Pseudotriton ruber (Sonnini de Manoncourt and Latreille, 1801) — Red Salamander
Proteidae Gray, 1825 — Mudpuppys and Olms
Necturus maculosus (Rafinesque, 1818) — Mudpuppy
Salamandridae Goldfuss, 1820 — Newts
Notophthalmus viridescens (Rafinesque, 1820) — Eastern Newt
Reptilia Laurenti, 1768 — Reptiles
Squamata (part) Oppel, 1811 — Snakes
Colubridae Oppel, 1811 — Harmless Egg-laying Snakes
Coluber constrictor Linnaeus, 1758 — North American Racer
Lampropeltis triangulum (Lacépède, 1789) — Eastern Milksnake
Opheodrys aestivus (Linnaeus, 1766) — Northern Rough Greensnake
Opheodrys vernalis (Harlan, 1827) — Smooth Greensnake
Pantherophis quadrivittatus (Holbrook, 1836) — Yellow Ratsnake
Crotalidae Oppel, 1811 — Pit Vipers
Agkistrodon contortrix (Linnaeus, 1766) — Eastern Copperhead
Crotalus horridus Linnaeus, 1758 — Timber Rattlesnake
Sistrurus catenatus (Rafinesque, 1818) — Eastern Massasauga
Dipsadidae Bonaparte, 1838 — Harmless Rear-Fanged Snakes
Carphophis amoenus (Say, 1825) — Common Wormsnake
Diadophis punctatus (Linnaeus, 1766) — Ring-necked Snake
Heterodon platirhinos Palisot de Beauvois in Sonnini de Manoncourt and Latreille, 1801 — Eastern Hog-nosed Snake
Natricidae Bonaparte, 1838 — Harmless Egg-Retaining Snakes
Clonophis kirtlandii (Kennicott, 1856) — Kirtland's Snake
Nerodia sipedon (Linnaeus, 1758) — Common Watersnake
Regina septemvittata (Say, 1825) — Queensnake
Storeria dekayi (Holbrook, 1839) — Dekay’s Brownsnake
Storeria occipitomaculata (Storer, 1839) — Red-bellied Snake
Thamnophis brachystoma (Cope, 1892) — Short-headed Gartersnake
Thamnophis saurita (Linnaeus, 1766) — Eastern Ribbonsnake
Thamnophis sirtalis (Linnaeus, 1758) — Common Gartersnake
Virginia valeriae Baird and Girard, 1853 — Smooth Earthsnake
Squamata (part) Oppel, 1811 — Other Lizards
Lacertidae Cope, 1864 — Lacertas and Wall Lizards
Podarcis siculus (Rafinesque, 1810) — Italian Wall Lizard   
Phrynosomatidae Fitzinger, 1843 — Sand and Spiny Lizards
Sceloporus undulatus (Bosc and Daudin in Sonnini de Manoncourt and Latreille, 1801) — Eastern Fence Lizard
Scincidae Gray, 1825 — Skinks
Plestiodon anthracinus (Baird, 1849) — Coal Skink
Plestiodon fasciatus (Linnaeus, 1758) — Common Five-lined Skink
Plestiodon laticeps (Schneider, 1801) — Broad-headed Skink
Testudines Batsch, 1788 — Turtles
Chelydridae Gray, 1870 — Snapping Turtles
Chelydra serpentina (Linnaeus, 1758) — Common Snapping Turtle
Emydidae Gray, 1825 — Box and Basking Turtles
Chrysemys picta (Schneider, 1783) — Painted Turtle
Clemmys guttata (Schneider, 1792) — Spotted Turtle
Emydoidea blandingii (Holbrook, 1838) — Blanding's Turtle
Glyptemys insculpta (LeConte, 1830) — Wood Turtle
Glyptemys muhlenbergii (Schoepff, 1801) — Bog Turtle
Graptemys geographica (LeSueur, 1817) — Northern Map Turtle
Malaclemys terrapin (Schoepff, 1793) — Diamondback Terrapin
Pseudemys rubriventris (LeConte, 1830) — Northern Red-bellied Cooter
Terrapene carolina Taylor, 1895 — Florida Box Turtle
Trachemys scripta (Thunberg in Schoepff, 1792) — Pond Slider
Kinosternidae Agassiz, 1857 — Mud and Musk Turtles
Kinosternon subrubrum (Bonnaterre, 1789) — Eastern Mud Turtle
Sternotherus odoratus (Latreille, 1802) — Eastern Musk Turtle
Trionychidae Bell, 1828 — Softshells
Apalone mutica (LeSueur, 1827) — Smooth Softshell
Apalone spinifera (LeSueur, 1827) — Spiny Softshell

(map source Wikipedia.com)

Pennsylvania Herpetological Publications
1901 Atkinson, D. A. The reptiles of Alleghany County, Pennsylvania. Annals of the Carnegie Museum 1():145-157
1906 Surface, H. A. The serpents of Pennsylvania. The Monthly Bulletin of the Division of Zoology, Pennsylvania State Department of Agriculture 4():Unpaginated
1906 Stone, W. Notes on reptiles and batrachians of Pennsylvania, New Jersey, and Delaware. American Naturalist 40():159-170
1908 Surface, H. A. First report on the lizards of Pennsylvania. Zoological Bulletin of the Division of Zoology of the Pennsylvania Department of Agriculture 5():236-258
1908 Surface, H. A. First report on the economy of Pennsylvania turtles. Zoological Bulletin of the Division of Zoology of the Pennsylvania Department of Agriculture 6():107-195
1928 Stewart, N. H. Some rare vertebrates of the Susquehanna Valley, Pennsylvania. Proceedings of the Pennsylvania Academy of Science 2():21-24
1930 Netting, M. Graham The occurrence of lizards in Pennsylvania. Annals of the Carnegie Museum 19(3):169-174
1930 Netting, M. Graham The poisonous snakes of Pennsylvania and the treatment of snakebites. Annals of the Carnegie Museum 19(3):175-184
1936 Netting, M. Graham Hand list of the amphibians and reptiles of Pennsylvania. Carnegie Mus. Pennsylvania Herpetology Leaflet 1():Unpaginated
1936 Netting, M. Graham The amphibians and reptiles of Indiana County, Pennsylvania. Proceedings of the Pennsylvania Academy of Science 10():25-28
1939 Pawling, R. O. The amphibians and reptiles of Union County, Pennsylvania. Herpetologica 1939(1):165-169
1940 Netting, M. Graham Handlist of the amphibians and reptiles of Pennsylvania. Biennial Report Pennsylvania Fish Commission for 1936-1938 1939():109-134
1941 Knepp, T. H. Some additions to the list of amphibians and reptiles of Bedford County. Proceedings of the Pennsylvania Academy of Science 15():164
1942 Conant, Roger Amphibians and reptiles from Dutch Mountain (Pennsylvania)and vicinity. American Midland Naturalist 27(1):154-170
1943 Baldauf, R. J. Handlist of Berks County amphibians and reptiles. Leaflets of the Mengel Natural History Society 1():1-8
1945 Smith, A. G. The amphibians and reptiles of Westmoreland County, Pennsylvania. Proceedings of the Pennsylvania Academy of Science 19():72-80
1946 Netting, M. Graham The amphibians and reptiles of Pennsylvania. Board Fish. Comm., Harrisburg, Pennsylvania. pp.
1948 Swanson, P. L. Notes on the amphibians of Venango County, Pennsylvania. American Midland Naturalist 40(2):362-371
1951 Heilman, R. A. A list of the amphibians and reptiles of Lebanon County, Pennsylvania. Proceedings of the Pennsylvania Academy of Science 25():44-46
1952 Swanson, P. L. The reptiles of Venango County, Pennsylvania. American Midland Naturalist 47():161-182
1954 Hudson, R. G. An annotated list of the reptiles and amphibians of the Unami Valley, Pennsylvania. Herpetologica 10():07-72
1955 Richmond, N. D. Poisonous snakes of Pennsylvania, how to recognise them. Carnegie Mus. Conservation Folder ():4 pp
1957 Bell, E. L. Some Herpetological records from Central Pennsylvania. Herpetologica 13():138-139
1959 Conant, Roger Lacerta colony still extant at Philadelphia. Copeia 1959():335-336
1960 Bell, E. L. Observations on the amphibians and reptiles of Huntingdon County, Pennsylvania. Proceedings of the Pennsylvania Academy of Science 34():159-161
1963 Stewart, G. D. ; Grant, R. R. County records for Pennsylvania and New Jersey. Bulletin of the Philadelphia Herpetological Society 11(3-4):57
1967 Ernst, C. H. Notes on the herpetofauna of the White Oak Bird Sanctuary. Bulletin of the Philadelphia Herpetological Society 15():32-36
1967 Feuer, R. C. Key to the adult amphibians and reptiles of Pennsylvania. Bulletin of the Philadelphia Herpetological Society 15():7-17
1974 McKinstry, D. M. ; Felege, S. Snakes of northwestern Pennsylvania. Bulletin of the Maryland Herpetological Society 10(1):29-31
1975 McKinstry, D. M. Notes on the herpetology of Presque Isle State Park, Erie, Pennsylvania. Bulletin of the Maryland Herpetological Society 11(1):21-26
1977 Groves, J. D. Priorities for effective management of the reptile fauna in Pennsylvania. Bulletin of the Philadelphia Herpetological Society 25():15-17
1977 Feuer, J. ; Feuer, R. C. Imperilled amphibians and reptiles of Pennsylvania and New Jersey. Bulletin of the Philadelphia Herpetological Society 25():26-28
1980 Steiner, R. P. Some effects of a controlled spring burn on vertebrate populations of the Blazing Star Prairie, Jennings Nature Reserve, Butler Co., Pa. (Abstract). Proceedings of the Pennsylvania Academy of Science 54(1):103
1981 Senecal Kearney, C. (ed. ) Pennsylvania species of special concern. Endangered Species Technical Bulletin 6(11):4-7
1982 McCoy, Clarence J. Amphibians and reptiles in Pennsylvania. Carnegie Museum of Natural History Special Publication 6():1-91
1982 Assetto, R. A list of reptiles and amphibians species collected randomly in lower Bucks County. Bulletin of the Philadelphia Herpetological Society 30():15-18
1985 McCoy, Clarence J. (ed. ) Species of special concern in Pennsylvania. Chapter 4 - amphibians and reptiles. Carnegie Museum of Natural History Special Publication 11():258-295
1985 Warny, P. R. A preliminary Herpetological survey in the Serpentine Barrens of Chester County, Pennsylvania. Bulletin of the Chicago Herpetological Society 20(1):23-24
1986 McCoy, Clarence J. New perspectives on threatened and endangered amphibians and reptiles in Pennsylvania. pp. 202-210. In: Majumdar, S.K.; Brenner, F.J. & Rhoads, A.F. (eds.). Endangered and threatened species programs in Pennsylvania and other states: Causes, issues and management. Pennsylvania Academy of Science 5():19
1986 Cypher, E. A. ; Yahner, R. H. ; Storm, G. L. ; Cypher, B. L. Flora and fauna survey in a proposed recreational area of Valley Forge National Historical Park. Proceedings of the Pennsylvania Academy of Science 60(1):47-50
1986 McCoy, Clarence J. Bibliography of Pennsylvanian herpetology: 1981-1986. Proceedings of the Pennsylvania Academy of Science 60(2):122-124
1986 Clements, D. June PHS field survey. Philadelphia Herpetological Society Newsletter 1986(2):
1989 McCoy, Clarence J. The amphibians and reptiles of Pennsylvania wetlands: An ecological survey. Pages in Pennsylvania Academy of Science Publications, Harrisburg, Pennsylvania. pp.
1990 Reinert, H. K. A profile and impact assessment of organized rattlesnake hunts in Pennsylvania. Journal of the Pennsylvania Academy of Science 64(3):136-144
1991 Brett, J. J. The Mountain and the migration: A guide to Hawk Mountain. Cornell University Press, Ithaca, New York. pp.
1991 Shaffer, L. L. and Clarence J. McCoy, Jr. Pennsylvania Amphibians and Reptiles. Pennsylvania Fish Commission, Harrisburg. pp.
1991 McKinstry, D. M. ; Lethaby, M. A. ; Cunningham, H. N. Amphibians and reptiles of Presque Isle State Park, Erie County, Pennsylvania. Journal of the Pennsylvania Academy of Science 65(1):17-23
1992 Allen, W. B. The snakes of Pennsylvania. Reptile & Amphibian Magazine, Pottsville, Pennsylvania. pp.
1992 Hulse, A. C. ; Hulse, K. L. New county records for amphibians and reptiles from Pennsylvania. Herpetological Review 23(2):62-64
1995 Bauer, A. M. ; Witwer, M. The herpetofauna of Nottingham Park and the Serpentine Barrens of Chester County, Pennsylvania. Bulletin of the Chicago Herpetological Society 30(1):3-9
1997 Lovich, J. E. Additional records of amphibians and reptiles from Indiana County, Pennsylvania. Journal of the Pennsylvania Academy of Science 71(1):35-38
1999 Shaffer, Larry L. Pennsylvania Amphibians and Reptiles. 3rd Revised edition. Pennsylvania Fish and Boat Commission, Bureau of Boating and Education, Educational Media Section, Harrisburg, Pennsylvania. 161pp.
2000 Harding, J. Reptiles and Amphibians of the Great Lakes Region. University of Michigan Press, Ann Arbor, Michigan. pp.
2001 Hulse, A. C. ; McCoy, Clarence J. ; Censky, E. J. Amphibians and reptiles of Pennsylvania and the northeast. Cornell University Press, Ithaca, New York. pp.
2001 Yahner, R. H. ; Piergallini, N. H. ; Ross, B. D. Survey of amphibians and reptiles in two types of managed forests in central Pennsylvania. Journal of the Pennsylvania Academy of Science 74(2-3):48-51
2001 Derge, K. L. ; Yahner, R. H. ; Mravintz, J. Survey of reptiles and amphibians in Gettysburg National Park with reference to Michaux State Forest. Journal of the Pennsylvania Academy of Science 75(2-3):43-49
2002 Gray, Brian S. Observations on the herpetofauna of Asbury Woods Greenway in Pennsylvania. Bulletin of the Chicago Herpetological Society 37(2):21-24
2006 Gray, Brian S. The amphibians and reptiles of the Asbury Woods Greenway, Erie County, Pennsylvania. Bulletin of the Maryland Herpetological Society 42(2):115-126
2006 Chambers, D. L. An updated list of the amphibians and reptiles of Indiana County, Pennsylvania (USA). Journal of the Pennsylvania Academy of Science 80(1):12-17
2007 Ross, B. ; Yahner, R. H. ; Karish, J. F. ; Kubel, J. E. Survey of amphibians and reptiles at Allegheny Portage Railroad National Historic Site and Johnstown Flood National Memorial, Pennsylvania. Journal of the Pennsylvania Academy of Science 81(1):3-7
2008 Meshaka, Walter E. ; Netting Huff, J. ; Leberman, R. C. Amphibians and reptiles of Powdermill Nature Reserve in western Pennsylvania. Journal of Kansas Herpetology 25():12-18
2008 Gray, Brian S. ; Lethaby, M. The amphibians and reptiles of Erie County, Pennsylvania. Bulletin of the Maryland Herpetological Society 44(2):49-69
2009 Gray, Brian S. Recent observations of the herpetofauna of a former national superfund site in Erie, Pennsylvania. Journal of Kansas Herpetology 31():9-11
2009 Meshaka, Walter E. ; Marshall, S. D. ; Guiher, T. J. ; Zemba, L. Grassland snake assemblages in central and western Pennsylvania and northeastern Ohio, USA. Herpetological Bulletin 110():8-19
2010 Steele, M. A. ; Brittingham, M. C. ; Maret, T. J. ; Merritt, J. F. (eds. ) Terrestrial vertebrates of Pennsylvania: A complete guide to species of conservation concern. Johns Hopkins University Press, Baltimore, Maryland. pp.
2010 Meshaka, Walter E. Seasonal activity and breeding seasons of snakes from Powdermill Nature Reserve in western Pennsylvania: the importance of site-specific data in land management programs. Herpetological Conservation and Biology 5(1):155-165
2010 Delis, P. R. ; Kindlin, C. ; Stewart, R. L. The herpetofauna of Letterkenny army depot, south-central Pennsylvania: A starting point to the long-term monitoring and management of amphibians and reptiles. Journal of Kansas Herpetology 34():11-16
2010 Meshaka, Walter E. Field observations of eleven species of snakes during twenty-six years at Hawk Mountain Sanctuary, Pennsylvania. Journal of Kansas Herpetology 36():17-20
2010 Stratford, J. ; Steele, M. A. Conservation of terrestrial vertebrates: An overview and synthesis Pages in Johns Hopkins University Press, Baltimore, Maryland. pp.
2010 Brittingham, M. C. ; Goodrich, L. J. Wildlife habitat: the key to abundance, distribution, and diversity. Pages in Johns Hopkins University Press, Baltimore, Maryland. pp.
2010 Williams, L. Pennsylvania's Wildlife Action Plan. Pages in Johns Hopkins University Press, Baltimore, Maryland. pp.
2010 Maret, T. J. The amphibians and reptiles. Pages in Johns Hopkins University Press, Baltimore, Maryland. pp.
2010 Steele, M. A. ; Reeder, D. M. ; Maret, T. J. ; Brittingham, M. C. Critical and emerging issues in the conservation of terrestrial vertebrates. Pages in Johns Hopkins University Press, Baltimore, Maryland. pp.
2014 Lovich, J. E. ; Ernst, C. H. ; Ernst, E. M. ; Riley, J. L. A 21-year study of seasonal and interspecific variation of hatchling emergence in a Nearctic freshwater turtle community: to overwinter or not to overwinter? Herpetological Monographs 28():93-109
2017 Hartzell, S. M. New township records for amphibians and reptiles in Columbia County, Pennsylvania, USA. Bulletin of the Chicago Herpetological Society 52(10):173-175
2018 Hughes, D. F. ; Delis, P. R. ; Meshaka, Walter E. Studies of a snake assemblage from artificial ponds at an activemilitary site in Pennsylvania. Herpetology Notes 11():35-48

THE CENTER FOR NORTH AMERICAN HERPETOLOGY — Accessed: Tuesday 28 January 2025 23:48 CT