1985
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Hillis, David M. Evolutionary genetics and systematics of New World frogs of the genus Rana: An analysis of ribosomal DNA, allozymes, and morphology. Dissertation. University of Kansas, Lawrence. pp.
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2014
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Feinberg, Jeremy A., Catherine E. Newman, Gregory J. Watkins-Colwell, Matthew D. Schlesinger, Brian Zarate, Brian R. Curry, H. Bradley Shaffer, and Joanna Burger. Cryptic diversity in metropolis: Confirmation of a new Leopard Frog species (Anura: Ranidae) from New York City and surrounding Atlantic Coast regions. PLoS One 9(10):1-15
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