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1920
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Boulenger, George A. A monograph of the American frogs of the genus Rana. Proceedings of the American Academy of Arts and Sciences (55):413-480
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1985
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Hillis, David M. Evolutionary genetics and systematics of New World frogs of the genus Rana: An analysis of ribosomal DNA, allozymes, and morphology. Dissertation. University of Kansas, Lawrence. pp.
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2014
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Feinberg, Jeremy A., Catherine E. Newman, Gregory J. Watkins-Colwell, Matthew D. Schlesinger, Brian Zarate, Brian R. Curry, H. Bradley Shaffer, and Joanna Burger. Cryptic diversity in metropolis: Confirmation of a new Leopard Frog species (Anura: Ranidae) from New York City and surrounding Atlantic Coast regions. PLoS One 9(10):1-15
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2018
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Schlesinger, Matthew D., Jeremy A. Feinberg, Nathan H. Nazdrowicz, J. D. Kleopfer, Jeffrey C. Beane, John F. Bunnell, Joanna Burger, Edward Corey, Kathy Gipe, Jesse W. Jaycox, Erik Kiviat, Jacob Kubel, Dennis P. Quinn, Christopher Raithel, Peter A. Scott, Sarah M. Wenner , Erin L. White, Brian Zarate, and H. Bradley Shaffer. Follow-up ecological studies for cryptic
species discoveries: Decrypting the leopard
frogs of the eastern U.S. PLoS ONE 13(11):e0205805
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2025
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Chambers, E. Anne, José Daniel Lara-Tufiño, Gustavo Campillo-García, Antonio Yolocalli Cisneros-Bernal, Daniel J. Dudek Jr., Virginia León-Règagnon, Josiah H. Townsend, Oscar Flores-Villela, and David M. Hillis. Distinguishing species boundaries from geographic variation. Proceedings of the National Academy of Sciences of the United States of America 122(19):e2423688122
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