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Baird, Spencer F. and Charles Girard. Characteristics of some new reptiles in the Museum of the Smithsonian Institution. Proceedings of the Academy of Natural Sciences, Philadelphia 6:68-70
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1963
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Lynn, R. T. Comparative behavior of the horned lizards, genus Phrynosoma, of the United States. Dissertation. University of Oklahoma, Norman, Oklahoma. 76pp.
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1965
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Lynn, R. T. A comparative study of the display behavior in Phrynosoma (Iguanidae). Southwestern Naturalist 10(1):25-30
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1987
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Montanucci, Richard R. A phylogenetic study of the horned lizards, genus Phrynosoma, based on skeletal and external morphology. Natural History Museum of Los Angeles County, Contributions in Science (390):1-36
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1987
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Clarke, Robert F. Second report of Phrynosoma modestum in Oklahoma. Bulletin of the Oklahoma Herpetological Society 12(1-4):16
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1993
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Wiens, John J. Phylogenetic relationships of phrynosomatid lizards and monophyly of the Sceloporus group. Copeia 1993(2):287-299
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1996
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Whiting, Martin J. and Dixon, James R. Phrynosoma modestum Catalogue of American Amphibians and Reptiles (630):1-6
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Montanucci, Richard R. Morphological variation in the gular fold in the horned lizard genus Phrynosoma (Iguania: Phrynosomatidae). Herpetologica 52(1):46-55
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2001
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Reeder, Tod W., and Richard R. Montanucci. Phylogenetic analysis of the horned lizards (Phrynosomatidae: Phrynosoma): Evidence from mitochondrial DNA and morphology. Copeia 2001(2):309-323
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2004
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Hodges, Wendy L. Evolution of viviparity in horned lizards (Phrynosoma): Testing the cold-climate hypothesis. Journal of Evolutionary Biology 17:1230–1237
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