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1852
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Baird, Spencer F. and Charles Girard. Descriptions of new species of reptiles, collected by the U. S. Exploring Expedition under the command of Capt. Charles Wilkes, U. S. N., First Part including the species from the western part of America. Proceedings of the Academy of Natural Sciences, Philadelphia 6:174-177
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1966
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Dumas, Philip C. Studies of the Rana species complex in the Pacific Northwest. Copeia 1966(1):60-74
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1977
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Brown, Herbert A. A case of interbreeding between Rana aurora and Bufo boreas (Amphibia, Anura). Journal of Herpetology 11(1):92-94
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1985
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Hillis, David M. Evolutionary genetics and systematics of New World frogs of the genus Rana: An analysis of ribosomal DNA, allozymes, and morphology. Dissertation. University of Kansas, Lawrence. pp.
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2001
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Macey, J. Robert, Jared L. Strasburg, Jennifer A. Brisson, Bance T. Vredenburg, Mark Jennings and Allan Larson. Molecular phylogenetics of western North American frogs of the Rana boylii species group. Molecular Phylogenetics and Evolution 19(1):131-143
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