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1928
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Pope, Clifford H. Some plethodontid salamanders from North Carolina and Kentucky with the description of a new race of Leurognathus. American Museum Novitates 306:1-19
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1993
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Arnold, Steven J., Nancy L. Reagan, and Paul A. Verrell. Reproductive isolation and speciation in plethodontid salamanders. Herpetologica 49(2):216-228
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1996
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Titus, Tom A. and Allan Larson. Molecular phylogenetics of Desmognathine salamanders (Caudata: Plethodontidae): A reevaluation of evolution in ecology, life history, and morphology. Systematic Biology 45(4):451-472
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2010
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Kozak, Kenneth H. and John J. Wiens. Accelerated rates of climatic-niche evolution underlie rapid species diversification. Ecology Letters 13:1378-1389
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2020
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Beamer, David A. and Trip Lamb. Towards rectifying limitations on species delineation in dusky salamanders (Desmognathus: Plethodontidae): An ecoregion-drainage sampling grid reveals additional cryptic clades. Zootaxa 4734(1):1–61
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2025
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Pyron, R. Alexander, Kyle A. O'connell, Edward A. Myers, David A. Beamer, and Hector Banos. Complex hybridization in a clade of polytypic salamanders (Plethodontidae: Desmognathus) uncovered by estimating higher-level phylogenetic networks. Systematic Biology 74(1):124-140
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