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1978
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Tilley, S. G., R. B. Merritt, B. Wu, and R. Highton. Genetic differentiation in salamanders of the Desmognathus ochrophaeus complex (Plethodontidae). Evolution 32():93–115
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2005
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Kozak, Kenneth H., Allan Larson, Ronald M. Bonett, and Luke J. Harmon. Phylogenetic analysis of ecomorphological civergence, community structure, and diversification rates in dusky salamanders (Plethodontidae: Desmognathus). Evolution 59(9):2000-2016
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2020
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Pyron, R. Alexander, K. A. O'Connell, Emily M. Lemmon, Alan R. Lemmon, and David A. Beamer. Phylogenomic data reveal reticulation and incongruence among mitochondrial candidate species in Dusky Salamanders (Desmognathus). Molecular Phylogenetics and Evolution 146(106751):1–13
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2020
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Beamer, David A. and Trip Lamb. Towards rectifying limitations on species delineation in dusky salamanders (Desmognathus: Plethodontidae): An ecoregion-drainage sampling grid reveals additional cryptic clades. Zootaxa 4734(1):1–61
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2022
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Pyron, R. Alexander and David A. Beamer. Systematics of the Ocoee Salamander (Plethodontidae: Desmognathus ocoee), with description of two new species from the southern Blue Ridge Mountains. Zootaxa (5190):207–240
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2022
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Pyron, R. Alexander, K. A. O'Connell, Emily M. Lemmon, Alan R. Lemmon, and David A. Beamer. Candidate-species delimitation in Desmognathus salamanders reveals gene flow across lineage boundaries, confounding phylogenetic estimation and clarifying hybrid zones. Ecology and Evolution 12(2: e8574):1–38
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