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Kansas Herpetofaunal Atlas

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Common Gartersnake

Thamnophis sirtalis (Linnaeus, 1758)

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Overview
HARMLESS. The pattern/color of this snake is variable. In general the Common Gartersnake is characterized by keeled scales, a single anal scale, a stripe on each side of the body situated on the second and third scale rows (counting from the belly up), absence of a crescent-shaped mark behind the angle of the jaw, and a pattern of two rows of alternating black spots on a between the stripes on the back. The belly is white, greenish, or gray, with a row of small dark spots along its edges. Adult males have longer tails than females, but females are larger than males.
The dorsal stripe may vary in the intensity of its coloration, but is usually well-defined and not encroached upon by adjacent lateral blotches. On specimens with orange or red dorsal stripes, the coloration usually starts with yellow or orange near the head and quickly transitions to the tip of the tail. The lateral stripes are less distinct and situated on second and third scale rows (counting from belly up) anteriorly. Between the stripes are two longitudinal rows of offset squarish black blotches producing a checkerboard pattern on a red (most wide-spread; parietalis type; 'red-sided' variant), yellow, green, or blue background. The belly is white, greenish, or gray, with a row of small dark spots along the edges.
Specimens from the extreme southwestern portion of their range in Kansas (Meade and Seward counties; annectens type; 'red-striped' variant) have a tan, black, yellow, or green alternating with the black blotches between the stripes. The dorsal stripe is bright red to orange. Some specimens from the Ozark Plateau of Cherokee County (sirtalis type; 'eastern' variant) (not noted by Hurd (1936)) have a muted pattern and color with an orange vertebral stripe and no lateral red markings.
The labials are heavily barred along their sutures and the posterior labials match the anterior labials in color. There may or may not be pair of light spots on top of the head along the suture of the parietal scales.
The young look like miniature adults with subdued coloration. Females have relatively larger heads.
Adults normally attain 41.0-71.0 cm (16-28 inches) in total length. The largest specimen from Kansas is a female (KU 189179) from Reno County with a total length of 113.0 cm (44½ inches) collected by George Ratzlaff on 5 March 1981. The maximum length throughout the range is 132.1 cm (52 inches) (Boundy, 1995; Powell et al., 2016). The maximum weight for a Kansas specimen is 410 grams (14½ ounces).

Distribution
Common throughout the eastern two-thirds of Kansas. Enters western Kansas along riparian corridors where it is seldom found far from them.Records mapped by Collins (1982) have since been reidentified (Hamilton County) or discredited/uncorroborated (Wallace County) (see Collins 1984)

Reproduction
Fitch and Maslin (1961) and Fitch (1965) studied this snake on the Great Plains, and much of the information from Kansas is based on their observations.
The Common Gartersnake is found in a wide variety of habitats, including marshes and wet meadows, margins of ponds, woodland and woodland edge, floodplains, and cultivated fields. It generally prefers areas with moderately moist vegetation. Platt (1985) found this reptile more common in the sand prairie habitat of western Harvey County than in the eastern part which lacked this habitat.
This hardy species is active from March to November at air temperatures ranging from 55° to 100°F. During warm periods from December to February, it frequently emerges from its underground winter retreats. Although this snake is active over a wide range of time and air temperatures, it appears to prefer air temperatures around 85°F.
Common Gartersnakes are active during the day and have a home range of 22- 35 acres. Estimates indicate a population density as high as three of these snakes per acre in northeastern Kansas (Fitch, 1982).
This snake mates during early spring and occasionally in the fall. On the first warm spring days, males search actively for females, locating them by scent. Several males may court the same female. The males lie alongside the female, their bodies rhythmically writhing. A successful male curls his tail beneath that of the passive female until his cloaca meets hers. Copulation occurs, and the female may drag the male along as she crawls.
Females give birth to young in late summer or fall. Litters are quite large in this species; the number of young per litter varies from four to 85 (Fitch, 1985), with an average of twenty. Collins (1974) recorded three females from Douglas County giving birth to 33, 35, and 35 young, respectively, during late July and August, the normal time of birth. Species of Thamnophis are viviparous and develop functional chorioallantoic and omphalallantoic placentae. Although embryos are predominantly lecithotrophic, substantial maternal transfer of respiratory gases, water, sodium, and calcium occurs, and experimental evidence demonstrates organic nutrient transfer consistent with incipient placentotrophy.
The Common Gartersnake feeds chiefly on frogs and earthworms but will also occasionally eat toads, small mice, and other small snakes. Henderson (1974) reported that examples of this snake from Douglas and Harvey counties fed almost exclusively on frogs.
Predators of this snake include hawks, large snakes, and mammals. Burt (1949) reported a young Common Gartersnake trapped and eaten by a large garden spider. Fitch (2001) reported the following prey items in decreasing abundance for northeast Kansas snakes: Allolbophora caliginosa (an earthworm), Lithobates blairi, Lithobates catesbeianus, Anaxyrus americanus, Anaxyrus woodhousii, Hyla chrysoscelis/versicolor, Acris crepitans, Pseudacris maculata, and three mammals (Microtus ochrogaster, Peromyscus leucopus, and Reithrodontomys megalotis).
Capron (1985) recorded 33 of these snakes killed by swathing (mowing) machines during agricultural harvests from May to mid-July in southern Kansas. Dillenbeck (1986) discovered nine of these harmless reptiles, all deliberately killed, along the Ninnescah River in southcentral Kansas.

Remarks
First reported in Kansas by Hallowell (1857) based on two specimens collected by Fort Riley surgeon William A. Hammond. The earliest available specimen (USNM 11934) was collected at Fort Riley by the post surgeon Bernard J. D. Irwin sometime during 1867.
Boundy (1999) found evidence to elevate the annectens phenotype present in north Texas and presumably (because he didn't include samples) those populations in Seward and southwest Meade counties in Kansas.
The annectens phenotype also shows up in extreme southwest Missouri and northwest Arkansas.
The annectens form was first mentioned from Kansas by Fitch (1961) and confirmed by Irwin (1992). The 'eastern' form was first mentioned from Kansas by Collins, Collins, and Taggart (2010).
Rundquist (1999) reported assumed hybridization with a captive male Thamnophis radix, however, given that long-term sperm storage and multiple paternity have been reported in T. sirtalis, there are other interpretations not considered.
Based on a captive specimen, Snider and Bowler (1992) reported a maximum longevity for this species of fourteen years.

References

Hallowell, Edward. 1856. Notice of a collection of reptiles from Kansas and Nebraska presented to the Academy of Natural Sciences, by Doctor Hammond, U. S. A. Proceedings of the Academy of Natural Sciences of Philadelphia 8:238-253.

Mozley, Annie E. 1877. List of Kansas snakes in the museum of the Kansas State University. Transactions of the Kansas Academy of Science 6:34-35.

Cragin, Francis W. 1880. A preliminary catalogue of Kansas reptiles and batrachians. Transactions of the Kansas Academy of Science 7:112-123.

Yarrow, Henry C. 1882. Check list of North American Reptilia and Batrachia with catalogue of specimens in U. S. National Museum. Bulletin of the United States National Museum (24):1-249.

Cragin, Francis W. 1885. Recent additions to the list of Kansas reptiles and batrachians, with further notes on species previously reported. Bulletin of the Washburn College Laboratory of Natural History 1(3):100-103.

Cragin, Francis W. 1885. Second contribution to the herpetology of Kansas, with observations on the Kansas fauna. Transactions of the Kansas Academy of Science 9:136-140.

Cope, Edward D. 1900. The crocodilians, lizards and snakes of North America. Pages 153-1270 in Report of the U. S. National Museum for the Year Ending June 30, 1898. , Washington, D. C.

Branson, Edwin B. 1904. Snakes of Kansas. University of Kansas Science Bulletin 2(13):353-430.

Ruthven, Alexander G. 1908. Variations and genetic relationships of the garter-snakes. Bulletin of the United States National Museum (61):1-201.

Hurter, Julius. 1911. Herpetology of Missouri. Transactions of the Academy of Science of St. Louis 20(5):59-274.

Linsdale, Jean M. 1925. Land Vertebrates of a Limited Area in Eastern Kansas. Thesis. University of Kansas, Lawrence, Kansas. 312 pp.

Burt, Charles E. 1927. An annotated list of the amphibians and reptiles of Riley County, Kansas. Occasional Papers of the Museum of Zoology, University of Michigan (189):12.

Linsdale, Jean M. 1927. Amphibians and reptiles of Doniphan County, Kansas. Copeia 1927(164):75-81.

Taylor, Edward H. 1929. A revised checklist of the snakes of Kansas. University of Kansas Science Bulletin 19(5):53-62.

Gloyd, Howard K. 1932. The herpetological fauna of the Pigeon Lake Region, Miami County, Kansas. Papers of the Michigan Academy of Science, Arts and Letters 15:389-408.

Burt, Charles E. 1933. Some distributional and ecological records of Kansas reptiles. Transactions of the Academy of Science of St. Louis 26:186-208.

Burt, Charles E. and W. L. Hoyle. 1934. Additional records of the reptiles of the central prairie region of the United States. Transactions of the Kansas Academy of Science 37:193-216.

Burt, Charles E. 1935. Further records of the ecology and distribution of amphibians and reptiles in the middle west. The American Midland Naturalist 16(3):311-366.

Hurd, Myron Alec. 1936. The reptiles of Cherokee County, Kansas. Thesis. Pittsburg State University, Pittsburg, Kansas. 103 pp.

Brumwell, Malcolm J. 1936. Distributional records of the reptilia and amphibians of Kansas. Privately printed. 22 pp.

Grant, Chapman. 1937. Herpetological notes from Central Kansas. The American Midland Naturalist 18(3):370-372.

Tihen, Joseph A. and James M. Sprague. 1939. Amphibians, reptiles, and mammals of the Meade County State Park. Transactions of the Kansas Academy of Science 42:499-512.

Hall, Henry H. and Hobart M. Smith. 1947. Selected records of reptiles and amphibians from southeastern Kansas. Transactions of the Kansas Academy of Science 49(4):447-454.

Brown, Bryce C. 1950. An Annotated Check List of the Reptiles and Amphibians of Texas. Baylor University Studies, Baylor, Texas.

Smith, Hobart M. 1950. Handbook of Amphibians and Reptiles of Kansas. University of Kansas, Museum of Natural History, Miscellaneous Publication (2):336.

Wolfenbarger, Keith. A. 1951. Systematic and Biological Studies on North American Chiggers of the genus Eutrombicula (Acarina, Trombiculidae). Thesis. University of Kansas, Lawrence. 77 pp.

Brumwell, Malcolm J. 1951. An ecological survey of the Fort Leavenworth Military Reservation. The American Midland Naturalist 45(1):187-231.

Schmidt, Karl P. 1953. A Check List of North American Amphibians and Reptiles. 6th Edition. University of Chicago Press, Chicago, Illinois. 280 pp.

Smith, Hobart M. 1956. Handbook of Amphibians and Reptiles of Kansas. Second edition. University of Kansas Museum of Natural History Miscellaneous Publication (9):1-356.

Loomis, Richard B. 1956. The chigger mites of Kansas (Acarina, Trombiculidae). University of Kansas Science Bulletin 37:1195-1443.

Steward, P. L. 1960. Lung-flukes of snakes, genera Thamnophis and Coluber, in Kansas. University of Kansas Science Bulletin 41:877-890.

Fitch, Henry S. and T. Paul Maslin. 1961. Occurrence of the garter snake, Thamnophis sirtalis in the Great Plains and Rocky Mountains. University of Kansas Publications Museum of Natural History 13(5):289-308.

Gish, Charles D. 1962. The Herpetofauna of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 34 pp.

Fitch, Henry S. 1965. An ecological study of the garter snake, Thamnophis sirtalis. University of Kansas Publications Museum of Natural History 15(10):493-504.

Choate, Jerry R. 1967. Wildlife in the Wakarusa Watershed of Northeastern Kansas. Kansas Biological Survey, Lawrence. 46 pp.

Fitch, Henry S. 1970. Reproductive cycles in lizards and snakes. University of Kansas Museum of Natural History Miscellaneous Publication (52):1-247.

Holman, J. Alan. 1972. Herpetofauna of the Kanopolis local fauna (Pleistocene: Yarmouth) of Kansas. Michigan Academician 5:87-98.

Henderson, Robert W. 1974. Resource partitioning among snakes of the University of Kansas Natural History Reservation: A preliminary analysis. Milwaukee Public Museum Contributions in Biology and Geology (1):1-11.

Collins, Joseph T. 1974. Amphibians and Reptiles in Kansas. University of Kansas Museum of Natural History Public Education Series (1):283 pp.

Rundquist, Eric M. 1975. Amphibians and Reptiles of Kingman County, Kansas. Privately Printed, Lawrence, Kansas. 3 pp.

Rundquist, Eric M. 1975. First KHS field trip yields three county records. Kansas Herpetological Society Newsletter (7):1-3.

Rundquist, Eric M. 1976. Field checklist (of) amphibians and reptiles of Kansas. Kansas Herpetological Society, Lawrence.

Caldwell, Janalee P. and Gregory. Glass. 1976. Vertebrates of the Woodson County State Fishing Lake and Game Management Area. Pages 62-76 in Preliminary inventory of the biota of Woodson County State Fishing Lake and Game Management Area. Report No. 5. State Biological Survey of Kansas, Lawrence.

Grow, David. 1976. Large garter snake caught at zoo. Kansas Herpetological Society Newsletter (12):8.

Capron, Marty B. and Jan Perry. 1976. A July weekend in Great Bend. Kansas Herpetological Society Newsletter (14):1-2.

Lardie, Richard L. 1976. Brood of Texas Garter Snakes from Oklahoma. Bulletin of the Oklahoma Herpetological Society 1(3):38.

Pisani, George R. 1976. Comments on the courtship and mating mechanics of Thamnophis (Reptilia, Serpentes, Colubridae). Journal of Herpetology 10(2):139-142.

Knight, James L. and Joseph T. Collins. 1977. The amphibians and reptiles of Cheyenne County, Kansas, Report Number 15. Kansas Biological Survey, Lawrence. 19 pp.

Trott, Gene. 1977. Chikaskia River wildlife study. Kansas Herpetological Society Newsletter (19):2-3.

Perry, Janice. 1977. KHS members achieve goal: Get Cottonmouth. Kansas Herpetological Society Newsletter (21):3-4.

Cink, Calvin L. 1977. Snake predation on Bell's Vireo nestlings. Wilson Bulletin 89(2):349-350.

Li, Amy. 1978. Comparative reproduction studies of two colubrid snakes (Thamnophis sirtalis parietalis, Lampropeltis triangulum syspila): Advanced Biology Report. Lawrence High School, Lawrence, Kansas. 15 pp.

Curl, Richard L. 1978. Final Environmental Statement: Milford Lake Kansas operation and maintenance. US Army Corps of Engineers, Kansas City District. 158 pp.

Fitch, Henry S. and E. Raymond Hall. 1978. A 20year record of succession on reseeded fields of tallgrass prairie on the Rockefeller Experimental Tract. University of Kansas Museum of Natural History Special Publication (4):1-15.

Li, Amy. 1978. Comparative reproduction studies of two colubrid snakes. Kansas Herpetological Society Newsletter (24):5-8.

Warner, M. and R. Wencel. 1978. Chikaskia River study held near Caldwell. Kansas Herpetological Society Newsletter (25):15-16.

Capron, Marty B. 1978. Four county collecting raid: A south central Kansas herping saga. Kansas Herpetological Society Newsletter (26):9-12.

Perry, Janice. 1978. KHS successful at Miami County State Lake. Kansas Herpetological Society Newsletter (27):5.

Collins, Joseph T. and Janalee P. Caldwell. 1978. New records of fishes, amphibians, and reptiles in Kansas for 1977. Technical Publication of the State Biological Survey of Kansas 6:70-88.

Gray, Peter and Eddie Stegall. 1979. A field trip to the Red Hills. Kansas Herpetological Society Newsletter (29):6-8.

Guarisco, Hank. 1979. Preliminary laboratory observations of predation by native Texas Garter Snakes upon hatchling Five-lined Skinks. Kansas Herpetological Society Newsletter (32):7-8.

Collins, Joseph T. 1979. New records of fishes, amphibians, and reptiles in Kansas for 1978. Technical Publication of the State Biological Survey of Kansas 8:56-66.

Clarke, Robert F. 1980. Herptiles and fishes of the western Arkansas River in Kansas. United States Army Corps of Engineers, Albuquerque, New Mexico. 55 pp.

Spencer, Dwight. 1980. Spencer, D. 1980. Ross Natural History Reservation: the first twenty years, 1959 to 1979. Emporia State University, Emporia, Kansas.. 64 pp.

Fitch, Henry S. 1980. Thamnophis sirtalis. Catalogue of American Amphibians and Reptiles (270):1-4.

Collins, Joseph T. 1980. New records of fishes, amphibians, and reptiles in Kansas for 1979. Technical Publication of the State Biological Survey of Kansas 9:1-11.

Collins, Joseph T. 1981. New records of fishes, amphibians, and reptiles in Kansas for 1980. Technical Publication of the State Biological Survey of Kansas 10:7-19.

Collins, Joseph T. 1982. Report to the Kansas Fish and Game Commission on the status of three amphibians in southeastern Kansas. Kansas Fish and Game Commission, Pratt. 57 pp.

Fitch, Henry S. 1982. Resources of a snake community in prairie-woodland habitat of northeastern Kansas. Pages 83-97 in Herpetological communities: A symposium of the Society for the Study of Amphibians and Reptiles and the Herpetologists League, August 1977.  Wildlife Research Reports 12. 239 pp. U. S. Fish and Wildlife Service, Washington, D. C.

Collins, Joseph T. 1982. Amphibians and Reptiles in Kansas. 2nd edition. University of Kansas Museum of Natural History Public Education Series (8).

Rogers, Karel L. 1982. Herpetofaunas of the Courland Canal and Hall Ash Local Faunas (Pleistoncene: Early Kansas) of Jewell Co., Kansas. Journal of Herpetology 16(2):174-177.

Overdeer, Danny C. 1983. Reptiles of the Mesquite Grasslands of Southwest Oklahoma. Dissertation. Oklahoma State University, Stillwater. 34 pp.

Miller, Larry L. 1983. Bourbon County field trip well attended and successful. Kansas Herpetological Society Newsletter (54):6-7.

Achen, Von, Pennie. H., and John L. Rakestraw. 1984. The role of chemoreception in the prey selection of neonate reptiles. Pages 163-172 in Vertebrate Ecology and Systematics— A Tribute to Henry S Fitch. Special Publication Number 10. The University of Kansas Museum of Natural History, Lawrence.

Holman, J. Alan. 1984. Herpetofaunas of the Duck Creek and Williams Local Faunas (Pleistocene: Illinoian) of Kansas. Pages 20-38 in Contributions in Quaternary Vertebrate Paleontology: A Volume in Memorial to John E. Guilday. Special Publication Number 8. Carnegie Museum of Natural History, Pittsburgh, Pennsylvania.

Eshelman, Ralph and Michael Hager. 1984. Two Irvingtonian (Medial Pleistocene) vertebrate faunas from northcentral Kansas. Pages 384-404 in Contributions in Quaternary Vertebrate Paleontology: A Volume in Memorial to John E. Guilday. Special Publication Number 8. Special Publication Number 8, Carnegie Museum of Natural History, Pittsburgh, Pennsylvania.

Brown, Kenneth L. 1984. Pomona: A plains village variant in eastern Kansas and western Missouri. Dissertation. University of Kansas, Lawrence, Kansas. 519 pp.

Heinrich, Mark L. 1984. Herpetofauna of the Konza Prairie Research Natural Area in the Flint Hills region of Kansas with respect to habitat selection. Thesis. Kansas State University, Manhattan, Kansas. 57 pp.

Secor, Stephen M. and Charles C. Carpenter. 1984. Distribution maps of Oklahoma reptiles. Oklahoma Herpetological Society Special Publication (3):1-57.

Collins, Joseph T. 1984. New records of fishes, amphibians, and reptiles in Kansas for 1983. Kansas Herpetological Society Newsletter (56):15-26.

Lynch, John D. 1985. Annotated checklist of the amphibians and reptiles of Nebraska. Transactions of the Nebraska Academy of Sciences 13:33-57.

Tanner, Wilmer W. 1985. Smales of western Chihuahua. Great Basin Naturalist 45(4):615-676.

Dillenbeck, Tom. 1986. Snake killings. Kansas Herpetological Society Newsletter (64):15.

Collins, Joseph T. 1986. New records of amphibians and reptiles in Kansas for 1986. Kansas Herpetological Society Newsletter (66):9-16.

Simmons, John E. 1987. September 1987 field trip report. Kansas Herpetological Society Newsletter (69):42894.

Holman, J. Alan. 1987. Snakes from the Robert Local Fauna (Late Wisconsinan) of Meade County, Kansas. Contributions from the Museum of Paleontology, University of Michigan 27(6):143-150.

Dillenbeck, Tom. 1988. Winter sightings. Kansas Herpetological Society Newsletter (71):12.

Miller, Larry L. 1988. Harper County KHS field trip well attended. Kansas Herpetological Society Newsletter (72):5-6.

Platt, Dwight R. 1989. Seasonal activity of snakes on a sand prairie. Pages 251-254 in Proceedings of the 11th North American Prairie Conference. University of Nebraska Press, Lincoln.

Collins, Joseph T. 1989. First Kansas herp counts held in 1989. Kansas Herpetological Society Newsletter (77):11-.

Simon, Martin P. and Joseph H. Dorlac. 1990. The results of a faunistic survey of reptiles and amphibians of Fort Leavenworth, Kansas. Kansas Department of Wildlife and Parks, Pratt. 11 pp.

Lardie, Richard L. 1990. Kansas threatened species and protection of the Gypsum Hills habitat. Kansas Herpetological Society Newsletter (80):14-15.

Collins, Joseph T. 1990. Results of second Kansas herp count held during April-May 1990. Kansas Herpetological Society Newsletter (81):10-12.

Collins, Joseph T. 1990. Maximum size records for Kansas amphibians and reptiles. Kansas Herpetological Society Newsletter (81):13-17.

Conant, Roger and Joseph T. Collins. 1991. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed. Houghton Mifflin Company, Boston, Massachusetts.

Fitch, Henry S. 1991. Reptiles and amphibians of the Kansas ecological reserves. Pages 71-74 in Ecology and Hydrology of Kansas Ecological Reserves and the Baker Wetlands. Multidisciplinary Guidebook 4. Kansas Academy of Science, Lawrence, Kansas.

Collins, Joseph T. 1991. New records of amphibians and reptiles in Kansas for 1990. Kansas Herpetological Society Newsletter (83):7-13.

Capron, Marty B. 1991. Unusual foraging behavior in water snakes (Nerodia) around drying pools in southcentral Kansas. Kansas Herpetological Society Newsletter (84):14-15.

Collins, Joseph T. 1991. Results of third Kansas herp count held during April-May 1991. Kansas Herpetological Society Newsletter (85):9-13.

Collins, Joseph T. 1992. New records of amphibians and reptiles in Kansas for 1991. Kansas Herpetological Society Newsletter (87):12-17.

Collins, Joseph T. 1992. Results of the fourth Kansas herp count held during April-May 1992. Kansas Herpetological Society Newsletter (89):10-.

Rundquist, Eric M. 1992. Results of the KHS 1992 fall field trip. Kansas Herpetological Society Newsletter (90):4.

Taggart, Travis W. 1992. Thamnophis sirtalis. Geographic distribution. Herpetological Review 23:92.

Irwin, Kelly J. 1992. Geographic distribution: Thamnophis sirtalis annectens. Herpetological Review 23(1):28.

Young, Eugene A. 1993. A Survey of the Vertebrates of Slate Creek Salt Marsh, Sumner County, Kansas, with an Emphasis on Waterbirds. Thesis. Fort Hays State University, Hays, Kansas. 189 pp.

Collins, Joseph T. and Suzanne L. Collins. 1993. Amphibians and Reptiles in Kansas. Third Edition. University Press of Kansas, Lawrence, Lawrence. 397 pp.

Collins, Joseph T. and Rundquist, Eric M. 1993. Results of the fifth Kansas herp count held during April-June 1993 . Kansas Herpetological Society Newsletter (94):7-11.

Viets, Brian E. 1993. An annotated list of the herpetofauna of the F. B., and Rena G. Ross Natural History Reservation. Transactions of the Kansas Academy of Science 96(1/2):103-113.

Fitch, Henry S. 1993. Relative abundance of snakes in Kansas. Transactions of the Kansas Academy of Science 96(3/4):213-224.

Fitzgerald, Eve C. and Charles Nilon. 1994. Classification of habitats for endangered and threatened species in Wyandotte County, Kansas. Kansas Department of Wildlife and Parks, Pratt, Kansas. 98 pp.

Riedle, J. Daren. 1994. Distribution of the Timber Rattlesnake (Crotalus horridus) in Chautauqua, Elk, and Montgomery counties, Kansas. Kansas Herpetological Society Newsletter (95):43051.

Collins, Joseph T. 1994. New records of amphibians and reptiles in Kansas for 1993. Kansas Herpetological Society Newsletter (97):15-19.

Rundquist, Eric M. 1994. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14.

Riedle, J. Daren. 1994. A survey of reptiles and amphibians at Montgomery County State Fishing Lake. Kansas Herpetological Society Newsletter (98):11-13.

Rundquist, Eric M. 1994. Additions and corrections [to the results of the sixth annual KHS herp counts held 1 April-31 May 1994]. Kansas Herpetological Society Newsletter (98):4.

Dloogatch, Michael A. (Editor). 1994. Herpetology 1994. Bulletin of the Chicago Herpetological Society 29(12):282-283.

Holman, J. Alan. 1995. Pleistocene Amphibians and Reptiles. Oxford University Press, New York. 243 pp.

Cink, Calvin L. 1995. Seasonal activity of snakes on a Kansas floodplain tallgrass prairie. Proceedings of the 14th North American Prairie Conference, Kansas State University, Manhattan :83-86.

Anderson, Lewis, Mark Shaw, Jeff Blodig, and Tom Walker. 1995. Report to the Kansas Department of Wildlife and Parks: Herps encountered during REmap project, summer 1994. Kansas Herpetological Society Newsletter (99):10-17.

Collins, Joseph T. 1995. New records of amphibians and reptiles in Kansas for 1994. Kansas Herpetological Society Newsletter (100):24-47.

Rundquist, Eric M. 1995. Results of the seventh annual KHS herp counts held 1 April-31 May 1995. Kansas Herpetological Society Newsletter (101):11-17.

Rundquist, Eric M. 1995. Additional KHS herp counts for 1995. Kansas Herpetological Society Newsletter (102):11-.

Rossman, Douglas A., Neil B. Ford, and Ricahrd A. Seigel. 1996. The Garter Snakes: Evolution and Ecology. University of Oklahoma Press, Norman. 332 pp.

Collins, Joseph T. 1996. New records of amphibians and reptiles in Kansas for 1995. Kansas Herpetological Society Newsletter (103):13-15.

Miller, Larry L. 1996. Results of the KHS 1995 fall field trip. Kansas Herpetological Society Newsletter (103):3.

Rundquist, Eric M. 1996. Results of the eighth annual KHS herp counts Held 1 April-31 May 1996. Kansas Herpetological Society Newsletter (104):6-17.

Rundquist, Eric M. 1996. Notes on the natural history of some Kansas amphibians and reptiles: Parasites. Kansas Herpetological Society Newsletter (105):16-17.

Miller, Larry L. 1996. Third graders conduct amphibian and reptile field study. Kansas Herpetological Society Newsletter (106):15.

Miller, Larry L. 1996. Many amphibian and reptile species identified during KHS 1996 fall field trip to Wabaunsee County. Kansas Herpetological Society Newsletter (106):2-3.

Rakestraw, J. 1996. Spring herp counts: A Kansas tradition. Reptile & Amphibian Magazine (March-April):75-80.

Rundquist, Eric M. 1997. Results of the ninth annual KHS herp counts held 1 April-31 May 1997. Kansas Herpetological Society Newsletter (108):12-17.

Rundquist, Eric M. 1997. Addendum to 1997 KHS herp counts. Kansas Herpetological Society Newsletter (109):14-15.

Collins, Joseph T. 1997. A report on the KHS fall field trip to the Marais des Cygnes wildlife refuges. Kansas Herpetological Society Newsletter (110):2-3.

Conant, Roger and Joseph T. Collins. 1998. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed, expanded. Houghton Mifflin Company, Boston, Massachusetts.

Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. 1998. A Key to Amphibians & Reptiles of the Continental United States and Canada. Univ Press of Kansas, Lawrence, Kansas. 131 pp.

Gamble, Jerre. 1998. Marais des Cygnes National Wildlife Refuge Comprehensive Conservation Plan. U.S. Fish and Wildlife Service, Hartford, Kansas. 91 pp.

Rundquist, Eric M. 1998. Results of the tenth annual KHS herp counts for 1998, held 1 April-31 May. Kansas Herpetological Society Newsletter (112):11-18.

Rundquist, Eric M. 1998. KDWP herp sting so far nets nine on Kansas and Federal charges. Kansas Herpetological Society Newsletter (112):5-6.

Collins, Joseph T. 1998. Results of the KHS silver anniversary fall field trip. Kansas Herpetological Society Newsletter (114):6-.

Boundy, Jeff. 1999. Systematics of the Common Garter Snake, Thamnophis sirtalis. Dissertation. Louisiana State University, Baton Rouge. 497 pp.

Rundquist, Eric M. 1999. Kansas Herpetological Society herp counts: A 10 year summary and evaluation. Kansas Herpetological Society Newsletter (115):42962.

Taggart, Travis W. 1999. Cherokee County fall 1999 herp count. Kansas Herpetological Society Newsletter (117):6.

Rundquist, Eric M. 1999. Garter snake hybridization. Kansas Herpetological Society Newsletter (118):15.

Fitch, Henry S. 2000. Population structure and biomass of some common snakes in central North America. Scientific Papers of the Natural History Museum, University of Kansas (17):1-7.

Miller, Larry L. 2000. A winter snake in Shawnee County. Kansas Herpetological Society Newsletter (119):11.

Taggart, Travis W. 2000. KHS spring field trip sets record for attendance. Kansas Herpetological Society Newsletter (120):5-5.

Taggart, Travis W. 2000. Biogeographic analysis of the reptiles (Squamata) in Ellis County, Kansas. Kansas Herpetological Society Newsletter (121):7-16.

Rundquist, Eric M. 2000. Results of the eleventh and twelfth annual KHS herpetofaunal counts for 1999-2000, held 1 April-31 May. Kansas Herpetological Society Newsletter (122):11-16.

Taggart, Travis W. 2000. Results of the KHS 2000 fall field trip. Kansas Herpetological Society Newsletter (122):6-8.

Johnson, Richard W. 2001. Spatial ecology of the Eastern Coachwhip (Masticophis flagellum flagellum) in and eastern Texas upland community. Thesis. Stephen F. Austin State University, Nacogdoches, Texas. 54 pp.

Fitch, Henry S. 2001. Further study of the garter snake, Thamnophis sirtalis, in northeastern Kansas. Scientific Papers of the Natural History Museum, University of Kansas (19):1-6.

Lardie, Richard L. 2001. The subspecific status of the Common Garter Snake, Thamnophis sirtalis, in western Oklahoma. Kansas Herpetological Society Newsletter (123):16-18.

Guarisco, Hank. 2001. Ode to an ophidion autumn. Kansas Herpetological Society Newsletter (123):19.

Taggart, Travis W. 2001. The KHS 2001 spring field trip: A rainy rendezvous. Kansas Herpetological Society Newsletter (124):12-14.

Ellis, Mark R. 2001. Reproduction in the Common Garter Snake in Shawnee County, Kansas. Kansas Herpetological Society Newsletter (125):12.

Rundquist, Eric M. 2001. Results of the thirteenth annual KHS herp counts for 2001, held 1 April-30 June. Kansas Herpetological Society Newsletter (125):13-16.

Kingsbury, Bruce and Joanna Gibson. 2002. Habitat Management Guidelines for Amphibians and Reptiles of the Midwest. Publication of Partners in Amphibian and Reptile Conservation, Address not given. 152 pp.

Fogell, Daniel D. 2002. Occurrence and relative abundance of amphibians and reptiles at Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. Interim Report. National Park Service, Washington, D.C.. 6 pp.

Riedle, J. Daren and A. Hynek. 2002. Amphibian and reptile inventory of the Kansas Army Ammunition Plant, Labette County, Kansas. Journal of Kansas Herpetology (2):18-20.

Ellis, Mark R. 2002. Fall 2002 KHS field trip to Washington County. Journal of Kansas Herpetology (2):4-5.

Taggart, Travis W. 2002. Results of the KHS 2002 fall field Trip. Journal of Kansas Herpetology (4):11-13.

Miller, Larry L. 2002. Osage County herp count II. Journal of Kansas Herpetology (4):15.

Miller, Larry L. 2002. Shawnee County herp count. Journal of Kansas Herpetology (4):15.

Miller, Larry L. 2002. Sumner County herp count. Journal of Kansas Herpetology (4):15.

Freeman, Craig C. 2003. A natural areas inventory of the Ft. Leavenworth Military Reservation, Leavenworth County, Kansas. II. Open-file Report No. 117. Kansas Biological Survey, Lawrence, Kansas. 199 pp.

Fogell, Daniel D. 2003. A herpetofaunal inventory of Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. National Park Service, Washington, D.C.. 59 pp.

Taggart, Travis W. 2003. KHS conducts first systematic road survey. Journal of Kansas Herpetology (6):11-12.

Platt, Dwight R. 2003. Lizards and snakes (Order Squamata) of Harvey County, Kansas. Journal of Kansas Herpetology (6):13-20.

Fitch, Henry S. 2003. Reproduction in snakes of the Fitch Natural History Reservation in northeastern Kansas. Journal of Kansas Herpetology (6):21-24.

Taggart, Travis W. 2003. Results of the 2003 KHS spring field trip to Wilson County. Journal of Kansas Herpetology (6):2-5.

Miller, Larry L. and Suzanne L. Miller. 2003. Wakarusa herp count. Journal of Kansas Herpetology (7):10.

Volkmann, Al. 2003. Cowley County herp count 1. Journal of Kansas Herpetology (7):7.

Collins, Joseph T. 2003. Douglas County herp count. Journal of Kansas Herpetology (7):8.

Miller, Larry L. 2003. Indian Creek herp count. Journal of Kansas Herpetology (7):9.

Suleiman, Gibran. 2003. Fort Riley herp count. Journal of Kansas Herpetology (7):9.

Taggart, Travis W. 2003. Results of the KHS 2003 fall field trip. Journal of Kansas Herpetology (8):14-15.

Fitch, Henry S., Scott Sharp, and Kylee Sharp. 2003. Snakes of the University of Kansas biotic succession area. Journal of Kansas Herpetology (8):20-21.

Setser, Kirk and John F. Cavitt. 2003. Effects of burning on snakes in Kansas, USA, tallgrass prairie. Natural Areas Journal 23(4):315-319.

Delisle, Jennifer M. and William H. Busby. 2004. Biological inventory for vertebrates at Fort Larned National Historic Site of the southern plains network. Natural Heritage Inventory, Kansas Biological Survey, Lawrence. 61 pp.

Fitch, Henry S. 2004. The effect of female size on number of eggs or young in snakes. Journal of Kansas Herpetology (9):11-12.

Fitch, Henry S. 2004. Food surplus and body size in local populations of snakes. Journal of Kansas Herpetology (10):14-16.

Volkmann, Al. 2004. Cowley County herp count. Journal of Kansas Herpetology (11):10.

Miller, Larry L. 2004. Sumner County herp count. Journal of Kansas Herpetology (11):11-12.

Ellis, Mark R. and Kathy Ellis. 2004. Wakarusa herp count. Journal of Kansas Herpetology (11):12.

Schmidt, Curtis J. 2004. Geographic distribution: Thamnophis sirtalis. Kansas. Journal of Kansas Herpetology (11):14.

Rehmeier, Ryan L. and Raymond S. Matlack. 2004. Life history. Thamnophis sirtalis. Diet. Journal of Kansas Herpetology (11):15.

Klug, Page. 2005. The effects of local grassland habitat and surrounding landscape compostion [sic] on the predators of grassland bird nests. Thesis. University of Nebraska at Omaha, Omaha, Nebraska. 143 pp.

Taggart, Travis W. 2005. Results of the KHS 2005 fall field trip [to Crawford County]. Journal of Kansas Herpetology (16):19-21.

Taggart, Travis W. 2006. Distribution and status of Kansas herpetofauna in need of information. State Wildlife Grant T7. Kansas Department of Wildlife and Parks, Pratt. vii + 106 pp.

Wilgers, Dustin J. and Eva A. Horne. 2006. Effects of different burn regimes on tallgrass prairie herpetofaunal species diversity and community composition in the Flint Hills, Kansas. Journal of Herpetology 40:73-84.

Wilgers, Dustin J., Eva A. Horne, Brett K. Sandercock, and Allan W. Volkmann. 2006. Effects of rangeland management on community dynamics of the herpetofauna of the tallgrass prairie. 62:378-388.

Wiens, Jonathan P. 2008. Habitat use and abundance of the Common Garter Snake, Thamnophis sirtalis, at the northern limit of its range in Manitoba. Thesis. University of Manitoba, Winnipeg, Canada. 132 pp.

Maxell, Bryce A., Paul Hendricks, M. T. Gates, and S. Lenard. 2009. Montana amphibian and reptile status assessment, literature review, and conservation plan. Montana Natural Heritage Program, University of Montana, Missoula, MT. 642 pp.

Murrow, Daniel G. 2009. KHS 2009 spring field trip. Journal of Kansas Herpetology (29):42769.

Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. 2010. Amphibians, Reptiles, and Turtles of Kansas. Eagle Mountain Publishing., Provo, Utah. 400 pp.

Murrow, Daniel G. 2010. Kansas Herpetological Society spring field trip. Journal of Kansas Herpetology (33):2-3.

Gray, Brian S. 2010. Distribution of native and exotic earthworms in the eastern United States: Implications for the ecology of vermivorous snakes. Bulletin of the Chicago Herpetological Society 45(5):73-86.

McMartin, David C. 2011. U. S. Army 2011 Fort Leavenworth Herpetofaunal Survey: 23 April - 09 May 2011. Privately printed, Leavenworth, Kansas. 33 pp.

Taggart, Travis W. 2011. Kansas Herpetological Society 2011 spring field trip to be held in Chautauqua County. Journal of Kansas Herpetology (37):5-7.

Taggart, Travis W. 2011. Results of the KHS Spring Field Trip to Chautauqua County. Journal of Kansas Herpetology (38):2-4.

Taggart, Travis W. 2011. Results of the Kansas Herpetological Society 2011 Fall Field Trip to Lovewell State Park. Journal of Kansas Herpetology (39):4-5.

McMartin, D. Chris. 2011. Herp Count: Fort Leavenworth Herpetofaunal Survey for 2011. Journal of Kansas Herpetology (39):8-9.

Houck, Mike. 2011. Fort Riley Herpetofaunal Survey for 2011. Journal of Kansas Herpetology (39):9.

Dloogatch, Michael A. (Editor). 2011. Herpetology 2011. Bulletin of the Chicago Herpetological Society 46(7):90.

Mooi, Randall D., Jonathan P. Wiens, and Gary S. Casper. 2011. Extreme color variation within populations Thamnophis sirtalis, in central North America, subspecies. Copeia 2011(2):187-200.

Rohweder, Megan R. 2012. Spatial conservation prioritization of Kansas for terrestrial vertebrates. Thesis. Fort Hays State University, Hays, Kansas. 151 pp.

Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. 2012. Key to the Herpetofauna of the Continental United States and Canada: Second Edition, Revised and Updated. Univ Press of Kansas, Lawrence, Kansas. 152 pp.

Kauffman, Greg Lee. 2013. Stable isotope analysis of a middle woodland population from north central Kansas. Thesis. University of Kansas, Lawrence, Kansas. 110 pp.

Miller, Larry L. 2013. Wellington Lake Herpetological Survey. Collinsorum 2(1/2):12.

Sinclair, Tom. 2013. A four-day spring snake count across northern Kansas. Collinsorum 2(1/2):9.

Taggart, Travis W. 2013. KHS 2012 Spring Field Trip to Bourbon County State Lake. Collinsorum 2(3/4):3.

Taggart, Travis W. 2013. KHS 2012 Fall Field Trip to Atchison County State Lake. Collinsorum 2(3/4):4.

Taggart, Travis W. 2013. KHS 2013 Spring Field Trip to Schermerhorn Park, Cherokee County. Collinsorum 2(3/4):4.

Taggart, Travis W. 2013. KHS 2013 Summer Field Trip to Coldwater Lake, Comanche County. Collinsorum 2(3/4):5.

Taggart, Travis W. 2013. KHS 2013 Fall Field Trip to Butler County State Lake. Collinsorum 2(3/4):6.

Mardis, Dexter and Kevin Scott. 2013. 2013 Kansas Herpetofaunal Counts. Collinsorum 2(3/4):7.

Johnson, Stephen R. 2013. Half-time herping on one big prairie. Bulletin of the Chicago Herpetological Society 48(5):65-66.

McMartin, D. Chris. 2014. Fort Leavenworth Heretofaunal Survey for 2013. Collinsorum 3(1):10.

Taggart, Travis W. 2014. Results of the 2014 KHS Fall Field Trip to Woodson County. Collinsorum 3(2-4):12.

Taggart, Travis W. 2014. Recent scientific and standard English name changes effecting the Kansas herpetofauna. Collinsorum 3(2-4):9-10.

Brown, Kasandra A. 2015. Occupancy Modeling Of Herpetofauna And Grassland Nesting Birds At Quivira National Wildlife Refuge. Thesis. Fort Hays State University, Hays, Kansas. 72 pp.

Houck, Mike. 2015. 2015 Fort Riley Herpetofaunal Count final report. Collinsorum 4(1):10-11.

Taggart, Travis W. 2015. Spring Field Trip to the Greenhorn Limestone of Russell County. Collinsorum 4(3):2.

Taggart, Travis W. 2015. Summer Field Trip In The Harvey County Sandhills. Collinsorum 4(3):3.

Taggart, Travis W. 2015. Fall Field Trip Held In Washington County. Collinsorum 4(3):4.

Gray, Brian S. 2015. Natural history observations of the Common Gartersnake, Thamnophis sirtalis, from northwest Pennsylvania. Bulletin of the Chicago Herpetological Society 50(5):61-72.

Pittman, Galen L., Henry S. Fitch, and W. Dean Kettle. 2016. Vertebrate animals on the Fitch Natural History Reservation (1948-2002). Kansas Biological Survey Report Number 188, Lawrence. 48 pp.

Powell, Robert, Roger Conant, and Joseph T. Collins. 2016. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. Houghton Mifflin Harcourt, Boston. 494 pp.

Taggart, Travis W. 2016. Spring 2016 KHS field trip to Clark County was a soggy success. Collinsorum 5(2-3):2-3.

Taggart, Travis W. 2016. Results of the KHS Summer field trip to Caney River, Chautauqua County, Kansas. Collinsorum 5(2-3):4-5.

Taggart, Travis W. 2016. Results of the KHS ‘Fall’ field trip to Barber County. Collinsorum 5(2-3):6-7.

Crother, Brian I. (editor). 2017. Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, with Comments Regarding Confidence in Our Understanding. Eighth edition. Herpetological Circulars (43):1-102.

Mardis, Dexter R. 2017. Results from three Herpetofaunal tallies at Wichita State University’s Youngmeyer Ranch in Northwestern Elk County. Collinsorum 6(1):8-10.

Wiese, Joshua D., Brice Krohn, and Andrew J. Caven. 2017. Common Gartersnake (Thamnophis sirtalis) mortality likely resulting from cold exposure following late winter hibernaculum emergence. Collinsorum 6(2-3).

Taggart, Travis W. 2017. Results of the 2017 KHS Spring Field Trip to Elk County, Kansas. Collinsorum 6(2-3):6-8.

Houck, Mike. 2018. Herp Count: Fort Riley Military Installation. Collinsorum 7(1):17.

Leao, Suelem Muniz, Eric R. Pianka, and Nicolás Pelegrin. 2018. Is there evidence for population regulation in amphibians and reptiles? Journal of Herpetology 52(1):28-33.

Rose, Jonathan P., Brian J. Halstead, Glenn D. Wylie, and Michael L. Casazza. 2018. Spatial and temporal variability in growth of Giant Gartersnakes: Plasticity, precipitation, and prey. Journal of Herpetology 52(1):40-49.

Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. 2019. Key to the Herpetofauna of the Continental United States and Canada. Third Edition. Univ Press of Kansas, Lawrence, Kansas. 192 pp.

Key, Kayla N., Banta, Joshua A., Gluesenkamp, Andrew G., Hertweck, Kate L., and Placyk, John S. 2019. Conservation biology of a southwestern endemic, the Texas Garter Snake, Thamnophis sirtalis annectens: An integrative perspective. The Southwestern Naturalist 64(1):8-19.

Daniel, Richard E. and Brian S. Edmond. 2020. Atlas of Missouri Amphibians and Reptiles for 2019. Privately printed, Columbia, Missouri. 86 pp.

Riedle, J. Daren. 2020. Revisiting Kansas Herpetological Society field trip and Herp Count data: Distributional patterns and trend data of Kansas amphibians and reptiles. Collinsorum 9(1):7-16.

Hullinger, Allison, Zackary Cordes, Daren Riedle, and William Stark. 2020. Habitat assessment of the Broad-headed Skink (Plestiodon laticeps) and the associated squamate community in eastern Kansas. Transactions of the Kansas Academy of Science 123(1-2):137-150.

Taggart, Travis W and Sarah L Taggart. 2021. Herp Count: Neosho County: KHS-2020-01. Collinsorum 9(3):11.

Taggart, Travis W and Sarah L Taggart. 2021. Herp Count: Cherokee County: KHS-2020-02. Collinsorum 9(3):11-12.

Riedle, J. Daren, Tamera D. Riedle, Zachary Riedle, and Greya Riedle. 2021. Herp Count: Pratt County: KHS-2020-05. Collinsorum 9(3):12.

Taggart, Travis W and Sarah L Taggart. 2021. Herp Count: Cherokee County: KHS-2020-03. Collinsorum 9(3):12.

Riedle, J. Daren, Tamera D. Riedle, Zachary Riedle, and Greya Riedle. 2021. Herp Count: Stafford County: KHS-2020-34. Collinsorum 9(3):16.

Hallas, Joshua M., Thomas L. Parchman, and Chris R. Feldman. 2022. Phylogenomic analyses resolve relationships among garter snakes (Thamnophis: Natricinae: Colubridae) and elucidate biogeographic history and morphological evolution. Molecular Phylogenetics and Evolution 167(2022):107374.

Russell, Elisabeth. 2023. Habitat associations and fine-scale movements of the Red-spotted Toad (Anaxyrus punctatus) in Kansas and the efficacy of remote telemetry for monitoring small-scale movements. Thesis. Fort Hays State University, Hays, Kansas. 81 pp.

Nuñez, Leroy P., Levi N Gray, David Weisrock, and Frank T Burbrink. 2023. The phylogenomic and biogeographic history of the Gartersnakes, Watersnakes, and Allies (Natricidae: Thamnophiini). Molecular Phylogenetics and Evolution 186(2023):107844.

Jones, Leonard N., II, Adam D. Leache, and Frank T. Burbrink. 2023. Biogeographic barriers and historic climate shape the phylogeography and demography of the Common Gartersnake. Journal of Biogeography 2012–2029:18.

Wright, Jake. 2024. Herpetofaunal assemblage at a former rock quarry in the Flint Hills of Kansas. Collinsorum 13(1):8-9.

Paysen, Joseph W. 2024. Common Gartersnake roadkill scavenging. Collinsorum 13(1):9-10.

Paysen, Joseph W., MacKenzie J. Smith, Carter Kruse, and Keith Geluso. 2024. Natural history notes. Thamnophis sirtalis. Diet. Herpetological Review 55(1):125.

Last Updated: 02/19/2026 8:05:15 AM CT

THE CENTER FOR NORTH AMERICAN HERPETOLOGY — Accessed: Tuesday 09 June 2026 13:11 CT