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Kansas Herpetofaunal Atlas

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Western Ribbonsnake

Thamnophis proximus (Say in James, 1822)

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Overview
HARMLESS. The Western Ribbonsnake is characterized by keeled scales, a single anal scale, a stripe on each side of the body situated on the third and fourth scale rows (counting from the belly up), upper lips without dark vertical bars, and a uniform olive-gray, dark green, or black skin color between the stripes. The stripe down the middle of the back is red, orange, or yellow and those on each side of the body are light yellow or cream. The belly is white, cream, or gray with some dark coloration along its edges. Females grow slightly larger than males.
Adults normally attain 51.0- 76.0 mm (20-30 inches) in total length. The largest specimen from Kansas is a female (FHSM 7243) from Edwards County with a total length of 101.3 cm (39½ inches) collected by James Gubanyi and Keith Coleman on 2 June 2002. The maximum length throughout the range is 126.7 cm (49­7⁄8 inches) (Boundy, 1995; Powell et al., 2016).

Distribution
Widespread in the eastern third of the state south of the Kansas River and largely confined to riparian corridors west of the Flint Hills.
Border vegetation around permanent bodies of water (swamps, marshes, ponds, rivers, ditches) and along the base of nearby rock outcrops where some individuals hibernate. Climbs onto piles of plants and low bushes.
Burt (1935) reported they are most abundant in extensive marshy areas that are filled with aquatic vegetation, such as reeds and cat-tails. They are often found in the reeds and grasses near shallow, meandering prairie streams as well, especially in the vicinity of those with sandy and gravelly bottoms.
Mozley (1877) first reported this species from Wallace County based on a specimen collected by Snow. This specimen no longer exists. Wilmer W. Tanner collected a specimen two miles east of Wallace (BYU 20742; no date is given). These are the westernmost reported specimens in Kansas, and additional corroborating specimens are desirable.

Reproduction
This slender, nervous snake frequents the edges of swamps, marshes, lakes, streams, and rivers. It is active from March to October and is both diurnal and nocturnal depending upon prevailing temperatures. This species spends the day foraging along water-edge vegetation or basking in the sun on matted reeds and grasses. When frightened, it glides swiftly across water to escape. Gloyd (1928) found this species active and feeding on frogs at night during April in Franklin County. During winter, it retreats to burrows underground to avoid cold temperatures.
Western Ribbonsnakes mate during April and May, but courtship is unknown. Each female gives birth to young anytime from late June to September. The number of young per litter ranges from 4 to 28 (Fitch, 1985), with an average of 12 or 13. Gloyd (1928) collected a female in Franklin County, which gave birth to eight young on 27 July. In Miami County, Gloyd (1932) found a female containing six well-developed young on 31 August. Collins (1974) obtained a gravid female collected in Franklin County, which died on 13 August and contained twelve well-developed young. Species of Thamnophis are viviparous and develop functional chorioallantoic and omphalallantoic placentae. Although embryos are predominantly lecithotrophic, substantial maternal transfer of respiratory gases, water, sodium, and calcium occurs, and experimental evidence demonstrates organic nutrient transfer consistent with incipient placentotrophy.
This species feeds on small frogs, toads, salamanders, and fish. Caldwell and Glass (1976) observed a Western Ribbonsnake eating a small Bullfrog in Woodson County. Taggart (1992) observed a Western Ribbonsnake consuming a Western Narrow-mouthed Toad in Chase County.
Predators of adult Western Ribbonsnakes include birds, mammals, and larger snakes. Young snakes are eaten by large frogs and fish (Collins, 1993).

Remarks
First reported in Kansas by Mozley (1877) based on a specimen collected by KU entomologist Francis A. Snow in Wallace County. The earliest existing specimen (KU 2068) was collected in Clark County in May of 1903.
In regions with extensive upland grasslands in the western two-thirds of the state, this taxon can be found well away from streams and ponds.
Based on a captive specimen, Snider and Bowler (1992) reported a maximum longevity for this snake of three years, seven months, and one day.

References

James, Edwin. 1822 "1823". Account of an expedition from Pittsburgh to the Rocky Mountains, performed in the years 1819 and ‘20, by order of the Hon. J. C. Calhoun, Sec’y of War: under the command of Major Stephen H. Long. From the notes of Major Long, Mr. T. Say, and other gentlemen of the exploring party. Compiled by Edwin James, botanist and geologist for the expedition. In two vols. – with an atlas. [Volume 1]. Henry Charles Carey and Isaac Lea, Philadelphia, Pennsylvania. 503 pp.

Mozley, Annie E. 1877. List of Kansas snakes in the museum of the Kansas State University. Transactions of the Kansas Academy of Science 6:34-35.

Cragin, Francis W. 1880. A preliminary catalogue of Kansas reptiles and batrachians. Transactions of the Kansas Academy of Science 7:112-123.

Yarrow, Henry C. 1882. Check list of North American Reptilia and Batrachia with catalogue of specimens in U. S. National Museum. Bulletin of the United States National Museum (24):1-249.

Garman, Samuel. 1884. The North American reptiles and batrachians. Bulletin of the Essex Institute 16:1-46.

Cragin, Francis W. 1885. Recent additions to the list of Kansas reptiles and batrachians, with further notes on species previously reported. Bulletin of the Washburn College Laboratory of Natural History 1(3):100-103.

Cragin, Francis W. 1885. Second contribution to the herpetology of Kansas, with observations on the Kansas fauna. Transactions of the Kansas Academy of Science 9:136-140.

Branson, Edwin B. 1904. Snakes of Kansas. University of Kansas Science Bulletin 2(13):353-430.

Ruthven, Alexander G. 1908. Variations and genetic relationships of the garter-snakes. Bulletin of the United States National Museum (61):1-201.

Linsdale, Jean M. 1925. Land Vertebrates of a Limited Area in Eastern Kansas. Thesis. University of Kansas, Lawrence, Kansas. 312 pp.

Linsdale, Jean M. 1927. Amphibians and reptiles of Doniphan County, Kansas. Copeia 1927(164):75-81.

Taylor, Edward H. 1929. A revised checklist of the snakes of Kansas. University of Kansas Science Bulletin 19(5):53-62.

Gloyd, Howard K. 1932. The herpetological fauna of the Pigeon Lake Region, Miami County, Kansas. Papers of the Michigan Academy of Science, Arts and Letters 15:389-408.

Burt, Charles E. 1933. Some distributional and ecological records of Kansas reptiles. Transactions of the Academy of Science of St. Louis 26:186-208.

Brennan, Lawrence A. 1934. A check list of the amphibians and reptiles of Ellis County, Kansas. Transactions of the Kansas Academy of Science 37:189-191.

Burt, Charles E. and W. L. Hoyle. 1934. Additional records of the reptiles of the central prairie region of the United States. Transactions of the Kansas Academy of Science 37:193-216.

Brennan, Lawrence A. 1935. Notes on the distribution of Amphibia and Reptilia of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 114 pp.

Burt, Charles E. 1935. Further records of the ecology and distribution of amphibians and reptiles in the middle west. The American Midland Naturalist 16(3):311-366.

Hurd, Myron Alec. 1936. The reptiles of Cherokee County, Kansas. Thesis. Pittsburg State University, Pittsburg, Kansas. 103 pp.

Brumwell, Malcolm J. 1936. Distributional records of the reptilia and amphibians of Kansas. Privately printed. 22 pp.

Grant, Chapman. 1937. Herpetological notes from Central Kansas. The American Midland Naturalist 18(3):370-372.

Brennan, Lawrence A. 1937. A study of the habitat of reptiles and amphibians of Ellis County, Kansas. Transactions of the Kansas Academy of Science 40:341-347.

Tihen, Joseph A. and James M. Sprague. 1939. Amphibians, reptiles, and mammals of the Meade County State Park. Transactions of the Kansas Academy of Science 42:499-512.

Marr, John C. 1944. Notes on amphibians and reptiles from the central United States. The American Midland Naturalist 32(2):478-490.

Hall, Henry H. and Hobart M. Smith. 1947. Selected records of reptiles and amphibians from southeastern Kansas. Transactions of the Kansas Academy of Science 49(4):447-454.

Smith, Hobart M. 1950. Handbook of Amphibians and Reptiles of Kansas. University of Kansas, Museum of Natural History, Miscellaneous Publication (2):336.

Wolfenbarger, Keith. A. 1951. Systematic and Biological Studies on North American Chiggers of the genus Eutrombicula (Acarina, Trombiculidae). Thesis. University of Kansas, Lawrence. 77 pp.

Brumwell, Malcolm J. 1951. An ecological survey of the Fort Leavenworth Military Reservation. The American Midland Naturalist 45(1):187-231.

Smith, Hobart M. 1956. Handbook of Amphibians and Reptiles of Kansas. Second edition. University of Kansas Museum of Natural History Miscellaneous Publication (9):1-356.

Loomis, Richard B. 1956. The chigger mites of Kansas (Acarina, Trombiculidae). University of Kansas Science Bulletin 37:1195-1443.

Steward, P. L. 1960. Lung-flukes of snakes, genera Thamnophis and Coluber, in Kansas. University of Kansas Science Bulletin 41:877-890.

Gish, Charles D. 1962. The Herpetofauna of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 34 pp.

Rossman, Douglas A. 1962. Thamnophis proximus (Say), a valid species of garter snake. Copeia 1962(4):741-748.

Rossman, Douglas A. 1963. The colubrid snake genus Thamnophis: A revision of the sauritus group. Bulletin of the Florida State Museum, Biological Sciences 7(3):99-178.

Choate, Jerry R. 1967. Wildlife in the Wakarusa Watershed of Northeastern Kansas. Kansas Biological Survey, Lawrence. 46 pp.

Brattstrom, Bayard H. 1967. A succession of Pliocene and Pleistocene snake fauna from the High Plains of the United States. Copeia 1967(1):188-202.

Rossman, Douglas A. 1970. Thamnophis proximus. Catalogue of American Amphibians and Reptiles (98):1-3.

Holman, J. Alan. 1972. Herpetofauna of the Kanopolis local fauna (Pleistocene: Yarmouth) of Kansas. Michigan Academician 5:87-98.

Collins, Joseph T. 1974. Amphibians and Reptiles in Kansas. University of Kansas Museum of Natural History Public Education Series (1):283 pp.

Rundquist, Eric M. 1975. Amphibians and Reptiles of Kingman County, Kansas. Privately Printed, Lawrence, Kansas. 3 pp.

Rundquist, Eric M. 1976. Field checklist (of) amphibians and reptiles of Kansas. Kansas Herpetological Society, Lawrence.

Caldwell, Janalee P. and Gregory. Glass. 1976. Vertebrates of the Woodson County State Fishing Lake and Game Management Area. Pages 62-76 in Preliminary inventory of the biota of Woodson County State Fishing Lake and Game Management Area. Report No. 5. State Biological Survey of Kansas, Lawrence.

Grow, David. 1976. The KHS goes to Chetopa. Kansas Herpetological Society Newsletter (13):2-3.

Miller, Larry L. 1976. KHS visits Elk County. Kansas Herpetological Society Newsletter (15):1-2.

Perry, Janice. 1977. KHS members achieve goal: Get Cottonmouth. Kansas Herpetological Society Newsletter (21):3-4.

Curl, Richard L. 1978. Final Environmental Statement: Milford Lake Kansas operation and maintenance. US Army Corps of Engineers, Kansas City District. 158 pp.

Perry, Janice. 1978. KHS successful at Miami County State Lake. Kansas Herpetological Society Newsletter (27):5.

Collins, Joseph T. and Janalee P. Caldwell. 1978. New records of fishes, amphibians, and reptiles in Kansas for 1977. Technical Publication of the State Biological Survey of Kansas 6:70-88.

Gray, Peter and Eddie Stegall. 1979. A field trip to the Red Hills. Kansas Herpetological Society Newsletter (29):6-8.

Clarke, Robert F. 1980. Herptiles and fishes of the western Arkansas River in Kansas. United States Army Corps of Engineers, Albuquerque, New Mexico. 55 pp.

Spencer, Dwight. 1980. Spencer, D. 1980. Ross Natural History Reservation: the first twenty years, 1959 to 1979. Emporia State University, Emporia, Kansas.. 64 pp.

Collins, Joseph T. 1980. New records of fishes, amphibians, and reptiles in Kansas for 1979. Technical Publication of the State Biological Survey of Kansas 9:1-11.

Collins, Joseph T. 1981. New records of fishes, amphibians, and reptiles in Kansas for 1980. Technical Publication of the State Biological Survey of Kansas 10:7-19.

Collins, Joseph T. 1982. Amphibians and Reptiles in Kansas. 2nd edition. University of Kansas Museum of Natural History Public Education Series (8).

Rogers, Karel L. 1982. Herpetofaunas of the Courland Canal and Hall Ash Local Faunas (Pleistoncene: Early Kansas) of Jewell Co., Kansas. Journal of Herpetology 16(2):174-177.

Simmons, John E. 1983. Fan fare. Kansas Herpetological Society Newsletter (54):16-18.

Collins, Joseph T. 1983. New records of fishes, amphibians, and reptiles in Kansas for 1982 . Technical Publication of the State Biological Survey of Kansas 13:9-21.

Holman, J. Alan. 1984. Herpetofaunas of the Duck Creek and Williams Local Faunas (Pleistocene: Illinoian) of Kansas. Pages 20-38 in Contributions in Quaternary Vertebrate Paleontology: A Volume in Memorial to John E. Guilday. Special Publication Number 8. Carnegie Museum of Natural History, Pittsburgh, Pennsylvania.

Eshelman, Ralph and Michael Hager. 1984. Two Irvingtonian (Medial Pleistocene) vertebrate faunas from northcentral Kansas. Pages 384-404 in Contributions in Quaternary Vertebrate Paleontology: A Volume in Memorial to John E. Guilday. Special Publication Number 8. Special Publication Number 8, Carnegie Museum of Natural History, Pittsburgh, Pennsylvania.

Brown, Kenneth L. 1984. Pomona: A plains village variant in eastern Kansas and western Missouri. Dissertation. University of Kansas, Lawrence, Kansas. 519 pp.

Heinrich, Mark L. 1984. Herpetofauna of the Konza Prairie Research Natural Area in the Flint Hills region of Kansas with respect to habitat selection. Thesis. Kansas State University, Manhattan, Kansas. 57 pp.

Secor, Stephen M. and Charles C. Carpenter. 1984. Distribution maps of Oklahoma reptiles. Oklahoma Herpetological Society Special Publication (3):1-57.

Collins, Joseph T. 1984. New records of fishes, amphibians, and reptiles in Kansas for 1983. Kansas Herpetological Society Newsletter (56):15-26.

Collins, Joseph T. 1984. New records of fishes, amphibians and reptiles in Kansas for 1984. Kansas Herpetological Society Newsletter (58):14-20.

Lynch, John D. 1985. Annotated checklist of the amphibians and reptiles of Nebraska. Transactions of the Nebraska Academy of Sciences 13:33-57.

Terry, P. A. 1986. Biological survey of the KS segments of Spring River and Shoal Creek. Part 1. Field Survey. Draft. Kansas Fish and Game, Pratt, Kansas. 67 pp.

Collins, Joseph T. 1986. New records of amphibians and reptiles in Kansas for 1985. Kansas Herpetological Society Newsletter (63):4.

Collins, Joseph T. 1986. New records of amphibians and reptiles in Kansas for 1986. Kansas Herpetological Society Newsletter (66):9-16.

Holman, J. Alan. 1986. Butler Spring herpetofauna of Kansas (Pleistocene: Illinoian) and its climatic significance. Journal of Herpetology 20(4):568-569.

Holman, J. Alan. 1987. Climatic significance of a late Illinoian herpetofauna from southwestern Kansas. Contributions from the Museum of Paleontology, University of Michigan 27(5):129-141.

Holman, J. Alan. 1987. Snakes from the Robert Local Fauna (Late Wisconsinan) of Meade County, Kansas. Contributions from the Museum of Paleontology, University of Michigan 27(6):143-150.

Collins, Joseph T. 1988. New records of amphibians and reptiles in Kansas for 1987. Kansas Herpetological Society Newsletter (71):13-19.

Miller, Larry L. 1988. Harper County KHS field trip well attended. Kansas Herpetological Society Newsletter (72):5-6.

Collins, Joseph T. 1989. First Kansas herp counts held in 1989. Kansas Herpetological Society Newsletter (77):11-.

Collins, Joseph T. 1989. New records of amphibians and reptiles in Kansas for 1989. Kansas Herpetological Society Newsletter (78):16-21.

Collins, Joseph T. 1990. Results of second Kansas herp count held during April-May 1990. Kansas Herpetological Society Newsletter (81):10-12.

Collins, Joseph T. 1990. Maximum size records for Kansas amphibians and reptiles. Kansas Herpetological Society Newsletter (81):13-17.

Conant, Roger and Joseph T. Collins. 1991. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed. Houghton Mifflin Company, Boston, Massachusetts.

Collins, Joseph T. 1991. New records of amphibians and reptiles in Kansas for 1990. Kansas Herpetological Society Newsletter (83):7-13.

Collins, Joseph T. 1991. Results of third Kansas herp count held during April-May 1991. Kansas Herpetological Society Newsletter (85):9-13.

Collins, Joseph T. 1992. New records of amphibians and reptiles in Kansas for 1991. Kansas Herpetological Society Newsletter (87):12-17.

Rundquist, Eric M. 1992. Results of the KHS 1992 fall field trip. Kansas Herpetological Society Newsletter (90):4.

Young, Eugene A. 1993. A Survey of the Vertebrates of Slate Creek Salt Marsh, Sumner County, Kansas, with an Emphasis on Waterbirds. Thesis. Fort Hays State University, Hays, Kansas. 189 pp.

Collins, Joseph T. and Suzanne L. Collins. 1993. Amphibians and Reptiles in Kansas. Third Edition. University Press of Kansas, Lawrence, Lawrence. 397 pp.

Viets, Brian E. 1993. An annotated list of the herpetofauna of the F. B., and Rena G. Ross Natural History Reservation. Transactions of the Kansas Academy of Science 96(1/2):103-113.

Fitch, Henry S. 1993. Relative abundance of snakes in Kansas. Transactions of the Kansas Academy of Science 96(3/4):213-224.

Collins, Joseph T. 1994. New records of amphibians and reptiles in Kansas for 1993. Kansas Herpetological Society Newsletter (97):15-19.

Rundquist, Eric M. 1994. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14.

Holman, J. Alan. 1995. Pleistocene Amphibians and Reptiles. Oxford University Press, New York. 243 pp.

Cink, Calvin L. 1995. Seasonal activity of snakes on a Kansas floodplain tallgrass prairie. Proceedings of the 14th North American Prairie Conference, Kansas State University, Manhattan :83-86.

Rundquist, Eric M. 1995. Results of the seventh annual KHS herp counts held 1 April-31 May 1995. Kansas Herpetological Society Newsletter (101):11-17.

Rundquist, Eric M. 1995. Additional KHS herp counts for 1995. Kansas Herpetological Society Newsletter (102):11-.

Rossman, Douglas A., Neil B. Ford, and Ricahrd A. Seigel. 1996. The Garter Snakes: Evolution and Ecology. University of Oklahoma Press, Norman. 332 pp.

Miller, Larry L. 1996. Results of the KHS 1995 fall field trip. Kansas Herpetological Society Newsletter (103):3.

Rundquist, Eric M. 1996. Results of the eighth annual KHS herp counts Held 1 April-31 May 1996. Kansas Herpetological Society Newsletter (104):6-17.

Miller, Larry L. 1996. Many amphibian and reptile species identified during KHS 1996 fall field trip to Wabaunsee County. Kansas Herpetological Society Newsletter (106):2-3.

Rakestraw, J. 1996. Spring herp counts: A Kansas tradition. Reptile & Amphibian Magazine (March-April):75-80.

Rundquist, Eric M. 1997. Results of the ninth annual KHS herp counts held 1 April-31 May 1997. Kansas Herpetological Society Newsletter (108):12-17.

Collins, Joseph T. 1997. A report on the KHS fall field trip to the Marais des Cygnes wildlife refuges. Kansas Herpetological Society Newsletter (110):2-3.

Conant, Roger and Joseph T. Collins. 1998. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed, expanded. Houghton Mifflin Company, Boston, Massachusetts.

Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. 1998. A Key to Amphibians & Reptiles of the Continental United States and Canada. Univ Press of Kansas, Lawrence, Kansas. 131 pp.

Gamble, Jerre. 1998. Marais des Cygnes National Wildlife Refuge Comprehensive Conservation Plan. U.S. Fish and Wildlife Service, Hartford, Kansas. 91 pp.

Rundquist, Eric M. 1998. Results of the tenth annual KHS herp counts for 1998, held 1 April-31 May. Kansas Herpetological Society Newsletter (112):11-18.

Rundquist, Eric M. 1999. Kansas Herpetological Society herp counts: A 10 year summary and evaluation. Kansas Herpetological Society Newsletter (115):42962.

Taggart, Travis W. 1999. Cherokee County fall 1999 herp count. Kansas Herpetological Society Newsletter (117):6.

Taggart, Travis W. 2000. KHS spring field trip sets record for attendance. Kansas Herpetological Society Newsletter (120):5-5.

Taggart, Travis W. 2000. Biogeographic analysis of the reptiles (Squamata) in Ellis County, Kansas. Kansas Herpetological Society Newsletter (121):7-16.

Rundquist, Eric M. 2000. Results of the eleventh and twelfth annual KHS herpetofaunal counts for 1999-2000, held 1 April-31 May. Kansas Herpetological Society Newsletter (122):11-16.

Taggart, Travis W. 2000. Results of the KHS 2000 fall field trip. Kansas Herpetological Society Newsletter (122):6-8.

Taggart, Travis W. 2001. The KHS 2001 spring field trip: A rainy rendezvous. Kansas Herpetological Society Newsletter (124):12-14.

Rundquist, Eric M. 2001. Results of the thirteenth annual KHS herp counts for 2001, held 1 April-30 June. Kansas Herpetological Society Newsletter (125):13-16.

Kingsbury, Bruce and Joanna Gibson. 2002. Habitat Management Guidelines for Amphibians and Reptiles of the Midwest. Publication of Partners in Amphibian and Reptile Conservation, Address not given. 152 pp.

Fogell, Daniel D. 2002. Occurrence and relative abundance of amphibians and reptiles at Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. Interim Report. National Park Service, Washington, D.C.. 6 pp.

Riedle, J. Daren and A. Hynek. 2002. Amphibian and reptile inventory of the Kansas Army Ammunition Plant, Labette County, Kansas. Journal of Kansas Herpetology (2):18-20.

Ellis, Mark R. 2002. Fall 2002 KHS field trip to Washington County. Journal of Kansas Herpetology (2):4-5.

Gubanyi, James E. and Keith Coleman. 2002. Size maxima: Thamnophis proximus. Journal of Kansas Herpetology (4):14.

Miller, Larry L. 2002. Sumner County herp count. Journal of Kansas Herpetology (4):15.

Freeman, Craig C. 2003. A natural areas inventory of the Ft. Leavenworth Military Reservation, Leavenworth County, Kansas. II. Open-file Report No. 117. Kansas Biological Survey, Lawrence, Kansas. 199 pp.

Fogell, Daniel D. 2003. A herpetofaunal inventory of Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. National Park Service, Washington, D.C.. 59 pp.

Suleiman, G. 2003. Fort Riley herpetofaunal count. Journal of Kansas Herpetology (5):11-12.

Collins, Joseph T. 2003. New records of amphibians and reptiles in Kansas for 2002. Journal of Kansas Herpetology (5):13-16.

Taggart, Travis W. 2003. Results of the 2003 KHS spring field trip to Wilson County. Journal of Kansas Herpetology (6):2-5.

Gubanyi, James E. 2003. Western Ribbon Snake reproduction. Journal of Kansas Herpetology (7):12.

Volkmann, Al. 2003. Cowley County herp count 1. Journal of Kansas Herpetology (7):7.

Taggart, Travis W. 2003. Results of the KHS 2003 fall field trip. Journal of Kansas Herpetology (8):14-15.

Delisle, Jennifer M. and William H. Busby. 2004. Biological inventory for vertebrates at Fort Larned National Historic Site of the southern plains network. Natural Heritage Inventory, Kansas Biological Survey, Lawrence. 61 pp.

Volkmann, Al. 2004. Cowley County herp count. Journal of Kansas Herpetology (11):10.

Murrow, Daniel G. 2004. Geographic distribution. Thamnophis proximus. Kansas. Journal of Kansas Herpetology (11):14.

Taggart, Travis W. 2005. Results of the KHS 2005 fall field trip [to Crawford County]. Journal of Kansas Herpetology (16):19-21.

Taggart, Travis W. 2006. Distribution and status of Kansas herpetofauna in need of information. State Wildlife Grant T7. Kansas Department of Wildlife and Parks, Pratt. vii + 106 pp.

Wilgers, Dustin J., Eva A. Horne, Brett K. Sandercock, and Allan W. Volkmann. 2006. Effects of rangeland management on community dynamics of the herpetofauna of the tallgrass prairie. 62:378-388.

Gomez, Nicholas J. 2008. Geographic distribution. Thamnophis proximus (Western Ribbon Snake). Journal of Kansas Herpetology (26):6.

Murrow, Daniel G. 2009. KHS 2009 spring field trip. Journal of Kansas Herpetology (29):42769.

Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. 2010. Amphibians, Reptiles, and Turtles of Kansas. Eagle Mountain Publishing., Provo, Utah. 400 pp.

Murrow, Daniel G. 2010. Kansas Herpetological Society spring field trip. Journal of Kansas Herpetology (33):2-3.

Taggart, Travis W. 2011. Kansas Herpetological Society 2011 spring field trip to be held in Chautauqua County. Journal of Kansas Herpetology (37):5-7.

Taggart, Travis W. 2011. Results of the KHS Spring Field Trip to Chautauqua County. Journal of Kansas Herpetology (38):2-4.

Taggart, Travis W. 2011. Results of the Kansas Herpetological Society 2011 Summer Field Trip to Scott State Park. Journal of Kansas Herpetology (39):2.

Rohweder, Megan R. 2012. Spatial conservation prioritization of Kansas for terrestrial vertebrates. Thesis. Fort Hays State University, Hays, Kansas. 151 pp.

Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. 2012. Key to the Herpetofauna of the Continental United States and Canada: Second Edition, Revised and Updated. Univ Press of Kansas, Lawrence, Kansas. 152 pp.

Taggart, Travis W. 2013. KHS 2012 Spring Field Trip to Bourbon County State Lake. Collinsorum 2(3/4):3.

Taggart, Travis W. 2013. KHS 2013 Spring Field Trip to Schermerhorn Park, Cherokee County. Collinsorum 2(3/4):4.

Mardis, Dexter and Kevin Scott. 2013. 2013 Kansas Herpetofaunal Counts. Collinsorum 2(3/4):7.

McMartin, D. Chris. 2014. Fort Leavenworth Heretofaunal Survey for 2013. Collinsorum 3(1):10.

Taggart, Travis W. 2014. Results of the 2014 KHS Spring Field Trip to Barber County. Collinsorum 3(2-4):11.

Taggart, Travis W. 2014. Results of the 2014 KHS Fall Field Trip to Woodson County. Collinsorum 3(2-4):12.

Taggart, Travis W. 2014. Recent scientific and standard English name changes effecting the Kansas herpetofauna. Collinsorum 3(2-4):9-10.

Taggart, Travis W. 2015. Spring Field Trip to the Greenhorn Limestone of Russell County. Collinsorum 4(3):2.

Taggart, Travis W. 2015. Summer Field Trip In The Harvey County Sandhills. Collinsorum 4(3):3.

Powell, Robert, Roger Conant, and Joseph T. Collins. 2016. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. Houghton Mifflin Harcourt, Boston. 494 pp.

Taggart, Travis W. 2016. Spring 2016 KHS field trip to Clark County was a soggy success. Collinsorum 5(2-3):2-3.

Taggart, Travis W. 2016. Results of the KHS Summer field trip to Caney River, Chautauqua County, Kansas. Collinsorum 5(2-3):4-5.

Taggart, Travis W. 2016. Results of the KHS ‘Fall’ field trip to Barber County. Collinsorum 5(2-3):6-7.

Crother, Brian I. (editor). 2017. Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, with Comments Regarding Confidence in Our Understanding. Eighth edition. Herpetological Circulars (43):1-102.

Mardis, Dexter R. 2017. Results from three Herpetofaunal tallies at Wichita State University’s Youngmeyer Ranch in Northwestern Elk County. Collinsorum 6(1):8-10.

Taggart, Travis W. 2017. Results of the 2017 KHS Spring Field Trip to Elk County, Kansas. Collinsorum 6(2-3):6-8.

Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. 2019. Key to the Herpetofauna of the Continental United States and Canada. Third Edition. Univ Press of Kansas, Lawrence, Kansas. 192 pp.

Daniel, Richard E. and Brian S. Edmond. 2020. Atlas of Missouri Amphibians and Reptiles for 2019. Privately printed, Columbia, Missouri. 86 pp.

Riedle, J. Daren. 2020. Revisiting Kansas Herpetological Society field trip and Herp Count data: Distributional patterns and trend data of Kansas amphibians and reptiles. Collinsorum 9(1):7-16.

Hullinger, Allison, Zackary Cordes, Daren Riedle, and William Stark. 2020. Habitat assessment of the Broad-headed Skink (Plestiodon laticeps) and the associated squamate community in eastern Kansas. Transactions of the Kansas Academy of Science 123(1-2):137-150.

Taggart, Travis W and Sarah L Taggart. 2021. Herp Count: Neosho County: KHS-2020-01. Collinsorum 9(3):11.

Mardis, Dexter R. 2021. Herp Count: Sumner County: KHS-2020-08. Collinsorum 9(3):12-13.

Taggart, Travis W. 2021. Herp Count: Russell County: KHS-2020-24. Collinsorum 9(3):14-15.

Taggart, Megan M. and Travis W. Taggart. 2021. Herp Count: Seward County: KHS-2020-27. Collinsorum 9(3):15.

Riedle, J. Daren, Tamera D. Riedle, Zachary Riedle, and Greya Riedle. 2021. Herp Count: Stafford County: KHS-2020-34. Collinsorum 9(3):16.

Riedle, J. Daren. 2021. Herp Count: Barton County: KHS-2020-28. Collinsorum 9(3):16.

Riedle, J. Daren. 2021. Herp Count: Pratt County: KHS-2020-33. Collinsorum 9(3):16.

Hallas, Joshua M., Thomas L. Parchman, and Chris R. Feldman. 2022. Phylogenomic analyses resolve relationships among garter snakes (Thamnophis: Natricinae: Colubridae) and elucidate biogeographic history and morphological evolution. Molecular Phylogenetics and Evolution 167(2022):107374.

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THE CENTER FOR NORTH AMERICAN HERPETOLOGY — Accessed: Tuesday 09 June 2026 13:09 CT