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Kansas Herpetofaunal Atlas

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Great Plains Ratsnake

Pantherophis emoryi (Baird and Girard, 1853)

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Overview
HARMLESS. The Great Plains Ratsnake is characterized by weakly keeled scales, a divided anal scale, a pattern of 25 -45 squarish brown blotches on its grayish back, and a checkerboard pattern of white and black or dark gray markings on its belly. Adult males have longer tails than females.
The young resemble adults and are similar in appearance to young Western Ratsnakes. The blotches on the back of the Great Plains Ratsnake are transverse, whereas the blotches on the Western Ratsnake are longitudinal.
Adults normally 61.0-122.0 cm (24-48 inches) in total length. The largest specimen from Kansas is a male (KU 192376) from Lyon County with a total length of 134.3 cm (52­5⁄8 inches) collected by H. A. Stephens on 15 May 1982. The maximum length throughout the range is 153 cm (60¼ inches) (Powell et al., 2016).

Distribution
This species occurs throughout most of Kansas, following the Arkansas, Smoky Hill, Solomon, Saline, Cimarron, and Republican rivers. It appears to be absent of much of the rest of the High Plains, the formerly glaciated regions north of the Kansas River and east of the Flint Hills, the Wellington Lowlands, and the lower Neosho River basin. There is an isolated population along the Bear Creek drainage in Stanton County, which is further supported by specimens in adjacent Colorado.

Reproduction
Great Plains Ratsnakes roam rocky hillsides and canyons and frequently inhabit caves in western Kansas. In eastern Kansas, they choose similar habitat in open woods or along woodland edge, avoiding heavily forested regions. This snake is active from March to September.
Great Plains Ratsnakes are primarily nocturnal, prowling for food. During the day, they remain hidden beneath rocks or in caves and crevices. Nothing is known of their home range or population density in Kansas. Heinrich and Kaufman (1985) observed 34 of these reptiles between 26 April and 26 September on the Konza Prairie near Manhattan; eleven had been killed by a controlled prairie fire on 26 April.
Using examples of the Great Plains Ratsnake from Kansas and Oklahoma, Gillingham (1979) described a three-phase courtship: the male chased the female ; then, jerking and writhing, he aligned his body with hers, and finally they copulated for 15 to 30 minutes. Number of eggs per clutch in this species probably ranges from three to thirty (Fitch, 1985), with an average of thirteen. Clark (1953) recorded two Kansas clutches of four and five eggs laid by this snake in early July. Mating probably occurs during spring, after emergence from winter inactivity.
This species constricts its prey and feeds primarily on small rodents and birds. Burt and Hoyle (1935) reported a specimen from a gypsum cave in Comanche County that was discovered hanging from slight irregularities on the roof.
Predators of the Great Plains Ratsnake include hawks, owls, mammals, and larger snakes.

Remarks
First reported from Kansas by Cope (1875) based on a specimen (no specific locality) in the collection at the United States National Museum. The earliest existing specimens (KU 2426-8) from Kansas were collected from Trego County in July of 1910. However, MCZ R5419 certainly predate KU 2426-8. They were received and cataloged in a lot of eight numbers from Francis W. Cragin collected near Manhattan and did not have a collected or received date associated with them. Other specimens cataloged just prior and after Cragin's series were dated 1879 to 1881.
USNM 2259 (collected at Hyatt, Anderson County by Samuel Arnyi) was published (p. 166) in Cochran (1961) as a paratype of Coluber rhinomegas Cope (a junior synonym of Pantherophis emoryi).
Noteworthy records (2 juvenile specimens) were collected in Stanton County (Taggart, 2006), and corroborate adjacent specimens in Colorado (Hammerson, 2001).
The epithet emoryi is in honor of Brigadier General William Hemsley Emory, who was chief surveyor of the U.S. Boundary Survey team of 1852 and collected specimens for the Smithsonian Institution. As such, it is sometimes referred to as Emory's Ratsnake.
When agitated, the Great Plains Ratsnake will shake its tail vigorously, which by itself makes no noise, but when it shakes among dry leaf litter, it can sound remarkably like a rattlesnake, and often leads to misidentification.
Burt (1935) reported on a den containing over a dozen of these snakes (young to adult) located on March 29, 1934, in a ravine 6 miles east of Winfield, Kansas, discovered on a warm day following a cold period. The den was at the base of a south slope in a clump of oak trees above a small stream. A perennial spring was situated nearby and much rock was in evidence. At one point a vertical rock was scaled to a height of about four feet and many snakes were revealed, including young Coluber constrictor and some adults of Diadophis punctatus in addition to Pantherophis emoryi.
Based on a captive specimen, Snider and Bowler (1992) reported a maximum longevity for this species of 21 years, one month, and 25 days.

References

Baird, Spencer F. and Charles Girard. 1853. Catalogue of North American Reptiles in the Museum of the Smithsonian Institution. Part 1. Serpents. Smithsonian Miscellaneous Collections 2(5):xvi + 172.

Kennicott, Robert. 1859. Notes on Coluber calligaster of Say, and a description of new species of serpents in the collection of the North Western University of Evanston, Illinois. Proceedings of the Academy of Natural Sciences of Philadelphia 11:98-100.

Cope, Edward D. 1875. Check-list of North American Batrachia and Reptilia; with a systematic list of the higher groups, and an essay on geographical distribution. Based on the specimens contained in the U.S. National Museum. Bulletin of the United States National Museum 1:1-104.

Jordan, David Starr. 1876. Manual of the Vertebrates of the Northern United States: Including the District East of the Mississippi River, and North of North Carolina and Tennessee, Exclusive of Marine Species. Jansen, McClurg, and Company, Chicago, Illinois.. 342 pp.

Jordan, David S. 1878. Manual of the Vertebrates of the Northern United States: Including the District East of the Mississippi River, and North of North Carolina and Tennessee, Exclusive of Marine Species. Second Edition, Revised, and Enlarged. Jansen, McClurg & Company, Chicago.

Cragin, Francis W. 1880. A preliminary catalogue of Kansas reptiles and batrachians. Transactions of the Kansas Academy of Science 7:112-123.

Davis, N. S. Jr. and Frank L. Rice. 1883. Descriptive catalogue of North American batrachia and reptilia, found east of Mississippi River. Illinois Natural History Survey Bulletin 1(5):71.

Cope, Edward D. 1900. The crocodilians, lizards and snakes of North America. Pages 153-1270 in Report of the U. S. National Museum for the Year Ending June 30, 1898. , Washington, D. C.

Brown, Arthur Erwin. 1901. A review of the genera and species of American snakes, north of Mexico. Proceedings of the Academy of Natural Sciences of Philadelphia 53(1):10-110.

Branson, Edwin B. 1904. Snakes of Kansas. University of Kansas Science Bulletin 2(13):353-430.

Ditmars, Raymond L. 1907. The Reptile Book; A comprehensive, Popularised Work on the Structure and Habits of the Turtles, Tortoises, Crocodilians, Lizards and Snakes which Inhabit the United States and Northern Mexico. Doubleday, Pae, and Company, New York. 472 pp.

Hurter, Julius and John K. Strecker, Jr. 1909. Amphibians and reptiles of Arkansas. Transactions of the Academy of Science of St. Louis 18(2):11-27.

Hurter, Julius. 1911. Herpetology of Missouri. Transactions of the Academy of Science of St. Louis 20(5):59-274.

Linsdale, Jean M. 1925. Land Vertebrates of a Limited Area in Eastern Kansas. Thesis. University of Kansas, Lawrence, Kansas. 312 pp.

Burt, Charles E. 1927. An annotated list of the amphibians and reptiles of Riley County, Kansas. Occasional Papers of the Museum of Zoology, University of Michigan (189):12.

Linsdale, Jean M. 1927. Amphibians and reptiles of Doniphan County, Kansas. Copeia 1927(164):75-81.

Taylor, Edward H. 1929. A revised checklist of the snakes of Kansas. University of Kansas Science Bulletin 19(5):53-62.

Gloyd, Howard K. 1932. The herpetological fauna of the Pigeon Lake Region, Miami County, Kansas. Papers of the Michigan Academy of Science, Arts and Letters 15:389-408.

Stejneger, Leonhard H. and Thomas Barbour. 1933. A Checklist of North American Amphibians and Reptiles. 3rd Edition. Harvard University Press, Cambridge, Massachusetts.

Burt, Charles E. 1933. Some distributional and ecological records of Kansas reptiles. Transactions of the Academy of Science of St. Louis 26:186-208.

Brennan, Lawrence A. 1934. A check list of the amphibians and reptiles of Ellis County, Kansas. Transactions of the Kansas Academy of Science 37:189-191.

Burt, Charles E. and W. L. Hoyle. 1934. Additional records of the reptiles of the central prairie region of the United States. Transactions of the Kansas Academy of Science 37:193-216.

Hibbard, Claude W. 1934. Notes on some cave bats of Kansas. Transactions of the Kansas Academy of Science 37:235-238.

Brennan, Lawrence A. 1935. Notes on the distribution of Amphibia and Reptilia of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 114 pp.

Burt, Charles E. 1935. Further records of the ecology and distribution of amphibians and reptiles in the middle west. The American Midland Naturalist 16(3):311-366.

Gloyd, Howard K. 1935. Some aberrant color patterns in snakes. Papers of the Michigan Academy of Science, Arts and Letters 20:661-668.

Hurd, Myron Alec. 1936. The reptiles of Cherokee County, Kansas. Thesis. Pittsburg State University, Pittsburg, Kansas. 103 pp.

Brumwell, Malcolm J. 1936. Distributional records of the reptilia and amphibians of Kansas. Privately printed. 22 pp.

Brennan, Lawrence A. 1937. A study of the habitat of reptiles and amphibians of Ellis County, Kansas. Transactions of the Kansas Academy of Science 40:341-347.

Schmidt, Karl Peterson and D. D. Davis. 1941. Field Book of Snakes of the United States and Canada. C.P. Putnam and Sons, New York. 365 pp.

Hudson, G. E. 1942. The amphibians and reptiles of Nebraska. Nebraska Conservation Bulletin 24:1-146.

Burt, Charles E. 1946. Description of Elaphe quivira, a new ratsnake from Kansas. Transactions of the Kansas Academy of Science 46(1):116.

Smith, Hobart M. 1950. Handbook of Amphibians and Reptiles of Kansas. University of Kansas, Museum of Natural History, Miscellaneous Publication (2):336.

Brumwell, Malcolm J. 1951. An ecological survey of the Fort Leavenworth Military Reservation. The American Midland Naturalist 45(1):187-231.

Dowling, Herndon G. 1952. A taxonomic study of the rat snakes, genus Elaphe Fitzinger. IV. A check list of the American forms. Occasional Papers of the Museum of Zoology, University of Michigan (541):1-12.

Schmidt, Karl P. 1953. A Check List of North American Amphibians and Reptiles. 6th Edition. University of Chicago Press, Chicago, Illinois. 280 pp.

Twente, John W., Jr. 1955. Aspects of a population study of cavern-dwelling bats. Journal of Mammalogy 36(3):379-390.

Smith, Hobart M. 1956. Handbook of Amphibians and Reptiles of Kansas. Second edition. University of Kansas Museum of Natural History Miscellaneous Publication (9):1-356.

Loomis, Richard B. 1956. The chigger mites of Kansas (Acarina, Trombiculidae). University of Kansas Science Bulletin 37:1195-1443.

Cochran, Doris M. 1961. Type specimens of reptiles and amphibians in the United States National Museum. Bulletin of the United States National Museum (220):1-289.

Gish, Charles D. 1962. The Herpetofauna of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 34 pp.

Choate, Jerry R. 1967. Wildlife in the Wakarusa Watershed of Northeastern Kansas. Kansas Biological Survey, Lawrence. 46 pp.

McLeran, V. 1973. Friendly constrictors. Kansas Fish and Game (30(2)):8-11.

Knight, James L., Eugene D. Fleharty, and Jerry D. Johnson. 1973. Noteworthy records of distribution and habits of some Kansas herptiles. Transactions of the Kansas Academy of Science 75(3):273-275.

Collins, Joseph T. 1974. Amphibians and Reptiles in Kansas. University of Kansas Museum of Natural History Public Education Series (1):283 pp.

Karns, Daryl, Ray E. Ashton, Jr., and Thomas Swearingen. 1974. Illustrated Guide to Amphibians and Reptiles in Kansas: An Identification Manual. University of Kansas Publications Museum of Natural History Public Education Series(2):viii + 18.

Thomas, Richard A. 1975. Taxonomic chaos: Elaphe guttata (Linnaeus), a case in point. Bulletin of the Maryland Herpetological Society 11(4):171-176.

Rundquist, Eric M. 1976. Field checklist (of) amphibians and reptiles of Kansas. Kansas Herpetological Society, Lawrence.

Caldwell, Janalee P. and Gregory. Glass. 1976. Vertebrates of the Woodson County State Fishing Lake and Game Management Area. Pages 62-76 in Preliminary inventory of the biota of Woodson County State Fishing Lake and Game Management Area. Report No. 5. State Biological Survey of Kansas, Lawrence.

Gillingham, James C. 1976. Comparative reproductive behavior of the rat snakes of Eastern North America, genus Elaphe. Dissertation. University of Oklahoma, Norman, Oklahoma. 68 pp.

Curl, Richard L. 1978. Final Environmental Statement: Milford Lake Kansas operation and maintenance. US Army Corps of Engineers, Kansas City District. 158 pp.

Capron, Marty B. 1978. Four county collecting raid: A south central Kansas herping saga. Kansas Herpetological Society Newsletter (26):9-12.

Collins, Joseph T. and Janalee P. Caldwell. 1978. New records of fishes, amphibians, and reptiles in Kansas for 1977. Technical Publication of the State Biological Survey of Kansas 6:70-88.

Clarke, Robert F. 1980. Herptiles and fishes of the western Arkansas River in Kansas. United States Army Corps of Engineers, Albuquerque, New Mexico. 55 pp.

Smith, Keith I. 1980. Observations of a Captive Kansas Least Weasel (Mustela nivalis). Thesis. Emporia State University, Emporia, Kansas. 55 pp.

Spencer, Dwight. 1980. Spencer, D. 1980. Ross Natural History Reservation: the first twenty years, 1959 to 1979. Emporia State University, Emporia, Kansas.. 64 pp.

Collins, Joseph T. 1980. New records of fishes, amphibians, and reptiles in Kansas for 1979. Technical Publication of the State Biological Survey of Kansas 9:1-11.

Collins, Joseph T. 1982. Report to the Kansas Fish and Game Commission on the status of three amphibians in southeastern Kansas. Kansas Fish and Game Commission, Pratt. 57 pp.

Fitch, Henry S. 1982. Resources of a snake community in prairie-woodland habitat of northeastern Kansas. Pages 83-97 in Herpetological communities: A symposium of the Society for the Study of Amphibians and Reptiles and the Herpetologists League, August 1977.  Wildlife Research Reports 12. 239 pp. U. S. Fish and Wildlife Service, Washington, D. C.

Collins, Joseph T. 1982. Amphibians and Reptiles in Kansas. 2nd edition. University of Kansas Museum of Natural History Public Education Series (8).

Collins, Joseph T. 1983. New records of fishes, amphibians, and reptiles in Kansas for 1982 . Technical Publication of the State Biological Survey of Kansas 13:9-21.

Brown, Kenneth L. 1984. Pomona: A plains village variant in eastern Kansas and western Missouri. Dissertation. University of Kansas, Lawrence, Kansas. 519 pp.

Heinrich, Mark L. 1984. Herpetofauna of the Konza Prairie Research Natural Area in the Flint Hills region of Kansas with respect to habitat selection. Thesis. Kansas State University, Manhattan, Kansas. 57 pp.

Secor, Stephen M. and Charles C. Carpenter. 1984. Distribution maps of Oklahoma reptiles. Oklahoma Herpetological Society Special Publication (3):1-57.

Collins, Joseph T. 1984. New records of fishes, amphibians, and reptiles in Kansas for 1983. Kansas Herpetological Society Newsletter (56):15-26.

Schwarting, Nancy. 1984. KHS field trip, May 1984. Kansas Herpetological Society Newsletter (57):3-4.

Miller, Larry L. 1985. KHS 1985 field trip to Kirwin Reservoir. Kansas Herpetological Society Newsletter (61):11-12.

Lynch, John D. 1985. Annotated checklist of the amphibians and reptiles of Nebraska. Transactions of the Nebraska Academy of Sciences 13:33-57.

Collins, Joseph T. 1986. New records of amphibians and reptiles in Kansas for 1985. Kansas Herpetological Society Newsletter (63):4.

Collins, Joseph T. 1986. New records of amphibians and reptiles in Kansas for 1986. Kansas Herpetological Society Newsletter (66):9-16.

Simmons, John E. 1987. September 1987 field trip report. Kansas Herpetological Society Newsletter (69):42894.

Collins, Joseph T. 1988. New records of amphibians and reptiles in Kansas for 1987. Kansas Herpetological Society Newsletter (71):13-19.

Collins, Joseph T. 1989. First Kansas herp counts held in 1989. Kansas Herpetological Society Newsletter (77):11-.

Kaufman, Glennis A. 1990. Population ecology, social organization, and mating systems in the Deer Mouse (Peromyscus maniculatus bairdii) in mixed-grass prairie in Kansas. Dissertation. Kansas State University, Manhattan, Kansas. 187 pp.

Collins, Joseph T. 1990. Results of second Kansas herp count held during April-May 1990. Kansas Herpetological Society Newsletter (81):10-12.

Collins, Joseph T. 1990. Maximum size records for Kansas amphibians and reptiles. Kansas Herpetological Society Newsletter (81):13-17.

Conant, Roger and Joseph T. Collins. 1991. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed. Houghton Mifflin Company, Boston, Massachusetts.

Fitch, Henry S. 1991. Reptiles and amphibians of the Kansas ecological reserves. Pages 71-74 in Ecology and Hydrology of Kansas Ecological Reserves and the Baker Wetlands. Multidisciplinary Guidebook 4. Kansas Academy of Science, Lawrence, Kansas.

Collins, Joseph T. 1991. New records of amphibians and reptiles in Kansas for 1990. Kansas Herpetological Society Newsletter (83):7-13.

Collins, Joseph T. 1991. Results of third Kansas herp count held during April-May 1991. Kansas Herpetological Society Newsletter (85):9-13.

Collins, Joseph T. 1992. New records of amphibians and reptiles in Kansas for 1991. Kansas Herpetological Society Newsletter (87):12-17.

Collins, Joseph T. 1992. Results of the fourth Kansas herp count held during April-May 1992. Kansas Herpetological Society Newsletter (89):10-.

Taggart, Travis W. 1992. Results of the KHS annual field trip to Sheridan County State Lake. Kansas Herpetological Society Newsletter (90):3-4.

Taggart, Travis W. 1992. KHS field trips. Kansas Herpetological Society Newsletter (91):3.

Collins, Joseph T. and Suzanne L. Collins. 1993. Amphibians and Reptiles in Kansas. Third Edition. University Press of Kansas, Lawrence, Lawrence. 397 pp.

Collins, Joseph T. and Rundquist, Eric M. 1993. Results of the fifth Kansas herp count held during April-June 1993 . Kansas Herpetological Society Newsletter (94):7-11.

Taggart, Travis W. 1993. Elaphe emoryi (Great Plains Rat Snake). USA: Kansas. Herpetological Review 24(2):67.

Viets, Brian E. 1993. An annotated list of the herpetofauna of the F. B., and Rena G. Ross Natural History Reservation. Transactions of the Kansas Academy of Science 96(1/2):103-113.

Fitch, Henry S. 1993. Relative abundance of snakes in Kansas. Transactions of the Kansas Academy of Science 96(3/4):213-224.

Anderson, Robert M. 1994. The effects of model transmitter implantation and surgery on feeding and movement in Elaphe guttata emoryi. Thesis. University of Nebraska at Omaha, Omaha, Nebraska. 45 pp.

Collins, Joseph T. 1994. New records of amphibians and reptiles in Kansas for 1993. Kansas Herpetological Society Newsletter (97):15-19.

Rundquist, Eric M. 1994. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14.

Rundquist, Eric M. 1994. Additions and corrections [to the results of the sixth annual KHS herp counts held 1 April-31 May 1994]. Kansas Herpetological Society Newsletter (98):4.

Dloogatch, Michael A. (Editor). 1994. Herpetology 1994. Bulletin of the Chicago Herpetological Society 29(12):282-283.

Smith, Hobart M., David Chiszar, J. R. Staley II, and K. Tepedelen. 1994. Populational relationships in the Corn Snake Elaphe guttata (Reptilia: Serpentes). Texas Journal of Science 46:259-292.

Adams, Steven P. 1995. Status of Bats in the Gypsum Hills of South-Central Kansas. Thesis. Fort Hays State University, Hays, Kansas. 39 pp.

Rundquist, Eric M. 1995. Results of the seventh annual KHS herp counts held 1 April-31 May 1995. Kansas Herpetological Society Newsletter (101):11-17.

Rundquist, Eric M. 1995. Additional KHS herp counts for 1995. Kansas Herpetological Society Newsletter (102):11-.

Bartlett, R. 1995. The captivating North American rat snakes. Reptiles 3(6):56-75.

Schultz, K. D. 1996. A Monograph of the Colubrid Snakes of the Genus Elaphe Fitzinger. Koeltz Scientific Books, Havlickuv Brod, Czech Republic.

Sparks, Dale W. 1996. Distribution, Natural History, Conservation Status, and Biogeography of Bats in Kansas. Thesis. Fort Hays State University, Hays, Kansas. 230 pp.

Miller, Larry L. 1996. Results of the KHS 1995 fall field trip. Kansas Herpetological Society Newsletter (103):3.

Rundquist, Eric M. 1996. Results of the eighth annual KHS herp counts Held 1 April-31 May 1996. Kansas Herpetological Society Newsletter (104):6-17.

Rakestraw, J. 1996. Spring herp counts: A Kansas tradition. Reptile & Amphibian Magazine (March-April):75-80.

Rundquist, Eric M. 1997. Results of the ninth annual KHS herp counts held 1 April-31 May 1997. Kansas Herpetological Society Newsletter (108):12-17.

Conant, Roger and Joseph T. Collins. 1998. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed, expanded. Houghton Mifflin Company, Boston, Massachusetts.

Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. 1998. A Key to Amphibians & Reptiles of the Continental United States and Canada. Univ Press of Kansas, Lawrence, Kansas. 131 pp.

Gamble, Jerre. 1998. Marais des Cygnes National Wildlife Refuge Comprehensive Conservation Plan. U.S. Fish and Wildlife Service, Hartford, Kansas. 91 pp.

Rundquist, Eric M. 1998. KDWP herp sting so far nets nine on Kansas and Federal charges. Kansas Herpetological Society Newsletter (112):5-6.

Rundquist, Eric M. 1999. Kansas Herpetological Society herp counts: A 10 year summary and evaluation. Kansas Herpetological Society Newsletter (115):42962.

Taggart, Travis W. 2000. KHS spring field trip sets record for attendance. Kansas Herpetological Society Newsletter (120):5-5.

Taggart, Travis W. 2000. Biogeographic analysis of the reptiles (Squamata) in Ellis County, Kansas. Kansas Herpetological Society Newsletter (121):7-16.

Rundquist, Eric M. 2000. Results of the eleventh and twelfth annual KHS herpetofaunal counts for 1999-2000, held 1 April-31 May. Kansas Herpetological Society Newsletter (122):11-16.

Taggart, Travis W. 2001. The KHS 2001 spring field trip: A rainy rendezvous. Kansas Herpetological Society Newsletter (124):12-14.

Rundquist, Eric M. 2001. Results of the thirteenth annual KHS herp counts for 2001, held 1 April-30 June. Kansas Herpetological Society Newsletter (125):13-16.

Kingsbury, Bruce and Joanna Gibson. 2002. Habitat Management Guidelines for Amphibians and Reptiles of the Midwest. Publication of Partners in Amphibian and Reptile Conservation, Address not given. 152 pp.

Fogell, Daniel D. 2002. Occurrence and relative abundance of amphibians and reptiles at Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. Interim Report. National Park Service, Washington, D.C.. 6 pp.

Collins, Joseph T. 2002. New records of amphibians and reptiles in Kansas for 2001. Journal of Kansas Herpetology (1):10-11.

Taggart, Travis W. 2002. Results of the KHS 2002 fall field Trip. Journal of Kansas Herpetology (4):11-13.

Utiger, Urs, Notker Helfenberger, Beat Schatti, Catherine Schmidt, Markus Ruf, and Vincent Ziswiler. 2002. Molecular systematics and phylogeny of Old and New World Ratsnakes, Elaphe AUCT., and related genera (Reptilia, Squamata, Colubridae). Russian Journal of Herpetology 9(2):105-124.

Burbrink, Frank T. 2002. Phylogeographic analysis of the Corn Snake (Elaphe guttata) complex as inferred from maximum likelihood and Bayesian analyses. Molecular Phylogenetics and Evolution 25:465-476.

Fogell, Daniel D. 2003. A herpetofaunal inventory of Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. National Park Service, Washington, D.C.. 59 pp.

Suleiman, G. 2003. Fort Riley herpetofaunal count. Journal of Kansas Herpetology (5):11-12.

Taggart, Travis W. 2003. Pantherophis emoryi winter activity. Journal of Kansas Herpetology (5):12.

Taggart, Travis W. 2003. Communal brumaculum. Journal of Kansas Herpetology (5):16.

Taggart, Travis W. 2003. KHS conducts first systematic road survey. Journal of Kansas Herpetology (6):11-12.

Taggart, Travis W. 2003. Results of the 2003 KHS spring field trip to Wilson County. Journal of Kansas Herpetology (6):2-5.

Lokke, John L. and Jill Lokke. 2003. Cowley County herp count 2. Journal of Kansas Herpetology (7):8.

Setser, Kirk and John F. Cavitt. 2003. Effects of burning on snakes in Kansas, USA, tallgrass prairie. Natural Areas Journal 23(4):315-319.

Lalley, M. L. 2004. Blood parasites of two species of rat snake (Elaphe spp.) from southeastern Nebraska and northeastern Kansas. Thesis. University of Nebraska at Omaha, Omaha, Nebraska.

Lalley, Maggie L. 2004. Blood parasites of two species of rat snake (Elaphe spp.) from southeastern Nebraska and northeastern Kansas. Thesis. University of Nebraska at Omaha, Omaha, Nebraska. 29 pp.

Delisle, Jennifer M. and William H. Busby. 2004. Biological inventory for vertebrates at Fort Larned National Historic Site of the southern plains network. Natural Heritage Inventory, Kansas Biological Survey, Lawrence. 61 pp.

Taggart, Travis W. 2004. Kansas Herpetological Society 2004 spring field trip. Journal of Kansas Herpetology (9):2.

Taggart, Travis W. 2004. Geographic distribution. Panthertherophis emoryi Kansas. Journal of Kansas Herpetology (11):14.

Taggart, Travis W. 2004. Results of the KHS 2004 fall feld trip . Journal of Kansas Herpetology (12):15-16.

Taggart, Travis W. 2005. Results of the KHS 2005 fall field trip [to Crawford County]. Journal of Kansas Herpetology (16):19-21.

Taggart, Travis W. 2006. Distribution and status of Kansas herpetofauna in need of information. State Wildlife Grant T7. Kansas Department of Wildlife and Parks, Pratt. vii + 106 pp.

Wilgers, Dustin J. and Eva A. Horne. 2006. Effects of different burn regimes on tallgrass prairie herpetofaunal species diversity and community composition in the Flint Hills, Kansas. Journal of Herpetology 40:73-84.

Wilgers, Dustin J., Eva A. Horne, Brett K. Sandercock, and Allan W. Volkmann. 2006. Effects of rangeland management on community dynamics of the herpetofauna of the tallgrass prairie. 62:378-388.

Taggart, Travis W., Joseph T. Collins, and Curtis J. Schmidt. 2007. Estimates of amphibian, reptile, and turtle mortality if Phostoxin is applied to 10,000 acres of prairie dog burrows in Logan County, Kansas. Kansas Department of Wildlife and Parks, Pratt. 5 pp.

Taggart, Travis W. 2008. KHS 2008 spring field trip. Journal of Kansas Herpetology (25):2-3.

Klug, Page E. 2009. Interactions between grassland birds and their snake predators: The potential for conservation conflicts in the tallgrass praire. Dissertation. Kansas State University, Manhattan, Kansas. 139 pp.

Murrow, Daniel G. 2009. KHS 2009 spring field trip. Journal of Kansas Herpetology (29):42769.

Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. 2010. Amphibians, Reptiles, and Turtles of Kansas. Eagle Mountain Publishing., Provo, Utah. 400 pp.

Murrow, Daniel G. 2010. Kansas Herpetological Society spring field trip. Journal of Kansas Herpetology (33):2-3.

Klug, Page E., Sara L. Jackerel, and Kimberly A. With. 2010. Linking snake habitat use to next predation risk in grassland birds: The dangers of shrub cover. Oecologia 162:803–813.

Taggart, Travis W. 2011. Kansas Herpetological Society 2011 spring field trip to be held in Chautauqua County. Journal of Kansas Herpetology (37):5-7.

Houck, Mike. 2011. Fort Riley Herpetofaunal Survey for 2011. Journal of Kansas Herpetology (39):9.

Klug, Page E., Jennifer Fill, and Kimberly A. With. 2011. Spatial ecology of Eastern Yellow-Bellied Racer (Coluber constrictor flaviventris) and Great Plains Rat Snake (Pantherophis emoryi) in a contiguous tallgrass-prairie landscape. Herpetologica 67(4):428-439.

Rohweder, Megan R. 2012. Spatial conservation prioritization of Kansas for terrestrial vertebrates. Thesis. Fort Hays State University, Hays, Kansas. 151 pp.

Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. 2012. Key to the Herpetofauna of the Continental United States and Canada: Second Edition, Revised and Updated. Univ Press of Kansas, Lawrence, Kansas. 152 pp.

Sinclair, Tom. 2013. A four-day spring snake count across northern Kansas. Collinsorum 2(1/2):9.

Baldwin, Mary Kate. 2013. “Herps in Havensville” Herp Surveys. Collinsorum 2(3/4):10.

Taggart, Travis W. 2013. KHS 2012 Spring Field Trip to Bourbon County State Lake. Collinsorum 2(3/4):3.

Taggart, Travis W. 2013. KHS 2013 Summer Field Trip to Coldwater Lake, Comanche County. Collinsorum 2(3/4):5.

Taggart, Travis W. 2014. Results of the 2014 KHS Spring Field Trip to Barber County. Collinsorum 3(2-4):11.

Taggart, Travis W. 2014. Recent scientific and standard English name changes effecting the Kansas herpetofauna. Collinsorum 3(2-4):9-10.

Houck, Mike. 2015. 2015 Fort Riley Herpetofaunal Count final report. Collinsorum 4(1):10-11.

Taggart, Travis W. 2015. Spring Field Trip to the Greenhorn Limestone of Russell County. Collinsorum 4(3):2.

Taggart, Travis W. 2015. Fall Field Trip Held In Washington County. Collinsorum 4(3):4.

Pittman, Galen L., Henry S. Fitch, and W. Dean Kettle. 2016. Vertebrate animals on the Fitch Natural History Reservation (1948-2002). Kansas Biological Survey Report Number 188, Lawrence. 48 pp.

Powell, Robert, Roger Conant, and Joseph T. Collins. 2016. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. Houghton Mifflin Harcourt, Boston. 494 pp.

Taggart, Travis W. 2016. Spring 2016 KHS field trip to Clark County was a soggy success. Collinsorum 5(2-3):2-3.

Taggart, Travis W. 2016. Results of the KHS ‘Fall’ field trip to Barber County. Collinsorum 5(2-3):6-7.

Taggart, Travis W. and J. Daren Riedle. 2017. A Pocket Guide to Kansas Amphibians, Turtles and Lizards. Great Plains Nature Center, Wichita, Kansas. 69 pp.

Crother, Brian I. (editor). 2017. Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, with Comments Regarding Confidence in Our Understanding. Eighth edition. Herpetological Circulars (43):1-102.

Mardis, Dexter R. 2017. Results from three Herpetofaunal tallies at Wichita State University’s Youngmeyer Ranch in Northwestern Elk County. Collinsorum 6(1):8-10.

Taggart, Travis W. 2017. Results of the 2017 KHS Spring Field Trip to Elk County, Kansas. Collinsorum 6(2-3):6-8.

Chen, Xin, Alan R. Lemmon, Emily Moriarty Lemmon, R. Alexander Pyron, and Frank T. Burbrink. 2017. Using phylogenomics to understand the link between biogeographic origins and regional diversification in ratsnakes. Molecular Phylogenetics and Evolution 111:206-218.

Meshaka, Walter E., Jr., and Curtis J. Schmidt. 2017. Reproductive traits in the Great Plains Ratsnake, Pantherophis emoryi (Baird and Girard, 1853), in northern Kansas. Transactions of the Kansas Academy of Science 120(3–4):170–174.

Houck, Mike. 2018. Herp Count: Fort Riley Military Installation. Collinsorum 7(1):17.

Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. 2019. Key to the Herpetofauna of the Continental United States and Canada. Third Edition. Univ Press of Kansas, Lawrence, Kansas. 192 pp.

Daniel, Richard E. and Brian S. Edmond. 2020. Atlas of Missouri Amphibians and Reptiles for 2019. Privately printed, Columbia, Missouri. 86 pp.

Riedle, J. Daren. 2020. Revisiting Kansas Herpetological Society field trip and Herp Count data: Distributional patterns and trend data of Kansas amphibians and reptiles. Collinsorum 9(1):7-16.

Myers, Edward A., McKelvy, Alexander D., and Burbrink, Frank T. 2020. Biogeographic barriers, Pleistocene refugia, and climatic gradients in the southeastern Nearctic drive diversification in Cornsnakes (Pantherophis guttatus complex). Molecular Ecology 29:797-811.

Marshall, Thomas L., E. Anne Chambers, Mikhail V. Matz, and David M. Hillis. 2021. How mitonuclear discordance and geographic variation have confounded species boundaries in a widely studied snake. Molecular Phylogenetics and Evolution (162):1-12.

Schmidt, Curtis J and Avery Schmidt. 2021. Herp Count: KHS-2020-4. Collinsorum 9(3):12.

Schmidt, Curtis J. 2021. Herp Count: Russell County: KHS-2020-19. Collinsorum 9(3):14.

Taggart, Meg, Amelia Jaeger, Jesse J. Taggart, and Travis W. Taggart. 2021. Herp Count: Ellis County: KHS-2020-21. Collinsorum 9(3):14.

Taggart, Travis W. 2021. Herp Count: Russell County: KHS-2020-24. Collinsorum 9(3):14-15.

Taggart, Travis W., Dan Fogell, and Christopher Visser. 2021. Herp Count: Russell County: KHS-2020-25. Collinsorum 9(3):15.

Burbrink, Frank and Sara Ruane. 2021. Contemporary philosophy and methods for studying speciation and delimiting species. Ichthyology & Herpetology 109(3):874-894.

Burbrink, Frank T., Edward A. Myers, R. Alexander Pyron. 2024. Understanding species limits through the formation of phylogeographic lineages. Ecology and Evolution 14(10):1-18.

Last Updated: 02/25/2024 10:15:17 PM CT

THE CENTER FOR NORTH AMERICAN HERPETOLOGY — Accessed: Tuesday 09 June 2026 13:15 CT