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Kansas Herpetofaunal Atlas

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Common Watersnake

Nerodia sipedon (Linnaeus, 1758)

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Overview
HARMLESS. The Common Watersnake is characterized by keeled scales, a divided anal scale, a dark brown, orange, yellow, red, and gray half-moon and speckled pattern widely spaced and scattered on the front part of a gray or whitish belly (these markings are closer together or fused on the rear portion of the belly, making it appear darker), and dark, complete bands on the front part of the body and alternating rows of dark blotches on the back and sides of the rear portion of the body. This snake may be gray or light brown, with dark gray or brown bands and blotches (young and young adults), or uniform dark gray with no pattern (old adults). Young specimens are more brightly colored and have a more contrasting pattern than adults. Adult males have longer tails than females. Adult females grow larger (twice the mass and 1.2 times the length) than males. Adults attain 56.0-106.7 cm (22-42 inches) in total length. The largest specimen from Kansas is a sex undetermined (though certainly a female) (KU 288637) from Jackson County with a total length of 120.8 cm (47½ inches) collected by James Gubanyi on 21 October 1998. The maximum length throughout the range is 149.9 cm (59 inches) (Conant and Collins, 1998). The maximum weight for a Kansas specimen is 480 grams (1 pound, 1 ounce).

Distribution
Largely confined to the riparian areas in the western half of Kansas, all the way into Colorado along the Arkansas and Republican rivers. In the eastern half, it is more widespread and may be commonly found around ponds and rocky streams.

Reproduction
This snake is found in almost any aquatic situation, from fast-flowing rocky streams and rivers to swamps, lakes, and marshes. It appears to become active at a lower temperature than most other snakes and has a preferred optimal temperature near 77°F. Taggart (1992) observed this snake sunning on the north bank of Shoal Creek in Cherokee County on 1 January, at an air temperature of 47°F; the creek had a thin layer of ice extending three to four feet from the shoreline.
The Common Watersnake is active from March to November, temperatures permitting. Like many snakes, it is active during the day in spring and fall and becomes more nocturnal in the summer. This species spends much of its time basking in the sun on branches and logs which overhang, or are near, water; it can be discovered easily by looking under rocks along streams in eastern Kansas.
Apparently, this snake spends the cold winter months in a variety of retreats, ranging from deep, dry crevices and holes on rocky, wooded hillsides to below water level in crayfish burrows in lowland areas.
Mating in this species occurs during the spring, and courtship involves the male positioning himself alongside the female and rubbing his chin on her neck while he spasmodically jerks his body to stimulate her to copulate. More than one male may copulate with a single female. The young are born from August to October. The number of young per litter varies from 6 to 66 (Fitch, 1985), with an average of 20-25. Some females may produce young only every other year. Pilch (1982) recorded a brood of 33 young born in early September to a female from Douglas County. Species of Nerodia are viviparous and develop functional placentae that support gas exchange and maternal–fetal transfer of water and ions. Stable-isotope evidence demonstrates maternal amino acid allocation to embryos during gestation, confirming organic nutrient transfer. Nevertheless, yolk remains the dominant source of embryonic dry mass; thus, Nerodia are best described as predominantly lecithotrophic viviparous snakes with incipient placentotrophy.
Diener (1957) showed that fish make up over 75 percent of the diet of the Common Watersnake. It also eats frogs and toads. Rundquist and Collins (1977) and Collins and Caldwell (1978) reported an adult Mudpuppy (KU 174546) discovered while being consumed by a Common Watersnake (Nerodia sipedon) in a shallow area of Shoal Creek at Schermerhorn Park, Cherokee County. Gray and Douglas (1989) recorded this snake preying on Southern Redbelly Dace in isolated pools on the Konza Prairie near Manhattan. The Common Watersnake uses an 'entrapment' behavior when preying on fish in drying pools. Riedle (1998) examined the stomach contents of three Common Watersnakes from Montgomery County. One adult snake, estimated to be 700 mm TL, was observed swallowing a 120 mm Channel Catfish (Ictalurus punctatus). A juvenile snake, approximately 207 mm TL, contained an adult Blanchard's Cricket Frog (Acris blanchardi), and another juvenile, approximately 210 mm TL, contained a small sunfish (Lepomis sp.), which was 35 mm in length.

Remarks
First reported in Kansas by Hallowell (1857) from specimens (no other associated data) received at the Academy of Natural Science, Philadelphia, from William A. Hammond, post surgeon at Fort Riley. Hallowell(1857) described Tropidonotus obliquus (= Nerodia sipedon) (ANSP 5016) from among those specimens. The earliest existing specimen (MCZ 5913) was collected in 1879 by J. Barbour. The MCZ catalog provides no other information, except that the specimen was received from the Smithsonian Institution in 1879 and that penciled-in notes question whether the actual locality is Kearney County, Kansas (where this species occurs along the Arkansas River and adjacent sand pits). This record is a bit problematic: while the listed state is Kansas, the locality is Fort Kearney (= Nebraska). At this time, the shipping point was often used as the locality for specimens collected up to 450 miles away. The Common Watersnake does occur along the Platte River near the site of Fort Kearney.
Beatson (1976) studied and analyzed the pattern and coloration of this snake in Chase County. He concluded that the change from the contrasting bright body color and pattern of young Common Watersnakes to the more uniform, darker pattern of old adults is related to changes in habitat and to pressure from predators. Young specimens usually inhabit small, rocky, fast-flowing streams, and their color and pattern blend readily with surrounding debris. Adults tend to migrate to larger, more open bodies of water where contrasting patterns and bright colors would be too conspicuous. The reason for this change in color pattern with age undoubtedly holds for the Plain-bellied Watersnake as well.
Based on a captive specimen, Snider and Bowler (1992) reported a maximum longevity for this snake of nine years, seven months, and 24 days.

References

Linné, Carl von (=Linneaus). 1758. Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species cum characteribus, differentiis, synonymis, locis. [System of Nature through the three kingdoms of nature, according to classes, orders, genera, species with characters, differences, synonyms, places.]. 10th Edition, Volume 1, L. Salvius, Stockholm. iv + 826 pp.

Hallowell, Edward. 1856. Notice of a collection of reptiles from Kansas and Nebraska presented to the Academy of Natural Sciences, by Doctor Hammond, U. S. A. Proceedings of the Academy of Natural Sciences of Philadelphia 8:238-253.

Mozley, Annie E. 1877. List of Kansas snakes in the museum of the Kansas State University. Transactions of the Kansas Academy of Science 6:34-35.

Cragin, Francis W. 1880. A preliminary catalogue of Kansas reptiles and batrachians. Transactions of the Kansas Academy of Science 7:112-123.

Yarrow, Henry C. 1882. Check list of North American Reptilia and Batrachia with catalogue of specimens in U. S. National Museum. Bulletin of the United States National Museum (24):1-249.

Cope, Edward D. 1900. The crocodilians, lizards and snakes of North America. Pages 153-1270 in Report of the U. S. National Museum for the Year Ending June 30, 1898. , Washington, D. C.

Brown, Arthur Erwin. 1901. A review of the genera and species of American snakes, north of Mexico. Proceedings of the Academy of Natural Sciences of Philadelphia 53(1):10-110.

Branson, Edwin B. 1904. Snakes of Kansas. University of Kansas Science Bulletin 2(13):353-430.

Ditmars, Raymond L. 1907. The Reptile Book; A comprehensive, Popularised Work on the Structure and Habits of the Turtles, Tortoises, Crocodilians, Lizards and Snakes which Inhabit the United States and Northern Mexico. Doubleday, Pae, and Company, New York. 472 pp.

Dyche, Lewis L. 1914. Enemies of fish. Pages 145-158 in Ponds, Pond Fish and Pond Fish Culture. State Department Fish and Game Bulletin No. 1, Kansas State Printing Office, Topeka.

Wooster, Lyman D. 1917. Nature Study Bulletin. Kansas State Printing Plant, Topeka, Kansas.. 63 pp.

Linsdale, Jean M. 1925. Land Vertebrates of a Limited Area in Eastern Kansas. Thesis. University of Kansas, Lawrence, Kansas. 312 pp.

Ortenburger, Arthur I. 1926. A report on the amphibians and reptiles of Oklahoma. Proceedings of the Oklahoma Academy of Science 6:89-100.

Burt, Charles E. 1927. An annotated list of the amphibians and reptiles of Riley County, Kansas. Occasional Papers of the Museum of Zoology, University of Michigan (189):12.

Linsdale, Jean M. 1927. Amphibians and reptiles of Doniphan County, Kansas. Copeia 1927(164):75-81.

Dolman, Katherine. 1929. Studies of Kansas Water Snakes. Thesis. University of Kansas, Lawrence. 69 pp.

Taylor, Edward H. 1929. A revised checklist of the snakes of Kansas. University of Kansas Science Bulletin 19(5):53-62.

Gloyd, Howard K. 1932. The herpetological fauna of the Pigeon Lake Region, Miami County, Kansas. Papers of the Michigan Academy of Science, Arts and Letters 15:389-408.

Burt, Charles E. 1933. Some distributional and ecological records of Kansas reptiles. Transactions of the Academy of Science of St. Louis 26:186-208.

Brennan, Lawrence A. 1934. A check list of the amphibians and reptiles of Ellis County, Kansas. Transactions of the Kansas Academy of Science 37:189-191.

Burt, Charles E. and W. L. Hoyle. 1934. Additional records of the reptiles of the central prairie region of the United States. Transactions of the Kansas Academy of Science 37:193-216.

Brennan, Lawrence A. 1935. Notes on the distribution of Amphibia and Reptilia of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 114 pp.

Burt, Charles E. 1935. Further records of the ecology and distribution of amphibians and reptiles in the middle west. The American Midland Naturalist 16(3):311-366.

Dunn, Emmet R. and Robert Allen. 1935. The redbellied watersnake in Pennsylvania. Copeia 1935(4):180-181.

Hurd, Myron Alec. 1936. The reptiles of Cherokee County, Kansas. Thesis. Pittsburg State University, Pittsburg, Kansas. 103 pp.

Brumwell, Malcolm J. 1936. Distributional records of the reptilia and amphibians of Kansas. Privately printed. 22 pp.

Grant, Chapman. 1937. Herpetological notes from Central Kansas. The American Midland Naturalist 18(3):370-372.

Brennan, Lawrence A. 1937. A study of the habitat of reptiles and amphibians of Ellis County, Kansas. Transactions of the Kansas Academy of Science 40:341-347.

Bond, Glenn Carl. 1938. Serological studies of the Reptilia. Dissertation. University of Kansas, Lawrence, Kansas. 110 pp.

Tihen, Joseph A. and James M. Sprague. 1939. Amphibians, reptiles, and mammals of the Meade County State Park. Transactions of the Kansas Academy of Science 42:499-512.

Schmidt, Karl Peterson and D. D. Davis. 1941. Field Book of Snakes of the United States and Canada. C.P. Putnam and Sons, New York. 365 pp.

Elliott, Alice. 1947. A preliminary survey and ecological study of the fishes of the South Ninnescah and Spring creek. Thesis. Kansas State University, Manhattan.

Hall, Henry H. and Hobart M. Smith. 1947. Selected records of reptiles and amphibians from southeastern Kansas. Transactions of the Kansas Academy of Science 49(4):447-454.

Smith, Hobart M. 1950. Handbook of Amphibians and Reptiles of Kansas. University of Kansas, Museum of Natural History, Miscellaneous Publication (2):336.

Brumwell, Malcolm J. 1951. An ecological survey of the Fort Leavenworth Military Reservation. The American Midland Naturalist 45(1):187-231.

Schmidt, Karl P. 1953. A Check List of North American Amphibians and Reptiles. 6th Edition. University of Chicago Press, Chicago, Illinois. 280 pp.

Smith, Hobart M. 1956. Handbook of Amphibians and Reptiles of Kansas. Second edition. University of Kansas Museum of Natural History Miscellaneous Publication (9):1-356.

Loomis, Richard B. 1956. The chigger mites of Kansas (Acarina, Trombiculidae). University of Kansas Science Bulletin 37:1195-1443.

Cliburn, J. William. 1957. Some southern races of the Common Water Snake, Natrix sipedon. Herpetologica 13:193-202.

Cliburn, J. W. 1960. The Phylogeny and Zoogeography of North American Natrix. Dissertation. University of Alabama, Tuscaloosa. 319 pp.

Nakmura, E. L., and H. M. Smith. 1960. A comparative study of selected characters in certain American species of watersnakes . Transactions of the Kansas Academy of Science 63(2):102-113.

Gish, Charles D. 1962. The Herpetofauna of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 34 pp.

Choate, Jerry R. 1967. Wildlife in the Wakarusa Watershed of Northeastern Kansas. Kansas Biological Survey, Lawrence. 46 pp.

Gier, Herschel T. 1967. Vertebrates of the Flint Hills. Transactions of the Kansas Academy of Science 70(1):51-59.

Brattstrom, Bayard H. 1967. A succession of Pliocene and Pleistocene snake fauna from the High Plains of the United States. Copeia 1967(1):188-202.

Holman, J. Alan. 1971. Herpetofauna of the sandahl local fauna (Pleistocene: Illinoian) of Kansas. Contributions from the Museum of Paleontology, University of Michigan 23(22):349-355.

Holman, J. Alan. 1972. Herpetofauna of the Kanopolis local fauna (Pleistocene: Yarmouth) of Kansas. Michigan Academician 5:87-98.

Eshelman, Ralph E. 1974. Geology and paleontology of the early Pleistocene Belleville Formation of north central Kansas. Dissertation. University of Michigan, Ann Arbor, Michigan. 137 pp.

Henderson, Robert W. 1974. Resource partitioning among snakes of the University of Kansas Natural History Reservation: A preliminary analysis. Milwaukee Public Museum Contributions in Biology and Geology (1):1-11.

Collins, Joseph T. 1974. Amphibians and Reptiles in Kansas. University of Kansas Museum of Natural History Public Education Series (1):283 pp.

Karns, Daryl, Ray E. Ashton, Jr., and Thomas Swearingen. 1974. Illustrated Guide to Amphibians and Reptiles in Kansas: An Identification Manual. University of Kansas Publications Museum of Natural History Public Education Series(2):viii + 18.

Rundquist, Eric M. 1975. Amphibians and Reptiles of Kingman County, Kansas. Privately Printed, Lawrence, Kansas. 3 pp.

Capron, Marty B. 1975. A trip through the Kansas Flint Hills. Kansas Herpetological Society Newsletter (8):4-5.

Eshelman, Ralph E. 1975. Geology and paleontology of the early Pleistocene (late Blancan) White Rock fauna from northcentral Kansas. University of Michigan Museum of Palenontology, Papers on Paleontology. (13):60.

Rundquist, Eric M. 1976. Field checklist (of) amphibians and reptiles of Kansas. Kansas Herpetological Society, Lawrence.

Caldwell, Janalee P. and Gregory. Glass. 1976. Vertebrates of the Woodson County State Fishing Lake and Game Management Area. Pages 62-76 in Preliminary inventory of the biota of Woodson County State Fishing Lake and Game Management Area. Report No. 5. State Biological Survey of Kansas, Lawrence.

Grow, David. 1976. The KHS goes to Chetopa. Kansas Herpetological Society Newsletter (13):2-3.

Capron, Marty B. and Jan Perry. 1976. A July weekend in Great Bend. Kansas Herpetological Society Newsletter (14):1-2.

Beatson, R. R. 1976. Environmental and genetical correlates of disruptive coloration in the water snake, Natrix s. sipedon. Evolution 30(2):241-252.

Rundquist, Eric M. and Joseph T. Collins. 1977. The amphibians of Cherokee County, Kansas. Kansas Biological Survey, Lawrence. 12 pp.

Knight, James L. and Joseph T. Collins. 1977. The amphibians and reptiles of Cheyenne County, Kansas, Report Number 15. Kansas Biological Survey, Lawrence. 19 pp.

Trott, Gene. 1977. Chikaskia River wildlife study. Kansas Herpetological Society Newsletter (19):2-3.

Perry, Janice. 1977. KHS members achieve goal: Get Cottonmouth. Kansas Herpetological Society Newsletter (21):3-4.

Curl, Richard L. 1978. Final Environmental Statement: Milford Lake Kansas operation and maintenance. US Army Corps of Engineers, Kansas City District. 158 pp.

Warner, M. and R. Wencel. 1978. Chikaskia River study held near Caldwell. Kansas Herpetological Society Newsletter (25):15-16.

Skie, Shelley and Martha Bickford. 1978. KHS takes to the field in July at Winfield. Kansas Herpetological Society Newsletter (26):42798.

Collins, Joseph T. and Janalee P. Caldwell. 1978. New records of fishes, amphibians, and reptiles in Kansas for 1977. Technical Publication of the State Biological Survey of Kansas 6:70-88.

Hibbard, Claude W., Richard J. Zakrzewski, Ralph E. Eshelman, Gordon Edmund, Clayton D. Griggs, and Caroline Griggs. 1978. Mammals from the Kanopolis Local Fauna, Pleistocene (Yarmouth) of Ellsworth County, Kansas. Contributions from the Museum of Paleontology, The University of Michigan 25(2):11-44.

Martin, Larry D. 1979. Survey of fossil vertebrates from east-central Kansas: Kansas River bank stabilization study. U.S. Army Corps of Engineers, Kansas City District. 55 pp.

Preston, Robert E. 1979. Late Pleistocene cold-blooded vertebrate faunas from the mid-continental United States, I. Reptilia: Testudines, Crocodilia. University of Michigan Museum of Palenontology, Papers on Paleontology. (19):1-53..

Gray, Peter and Eddie Stegall. 1979. A field trip to the Red Hills. Kansas Herpetological Society Newsletter (29):6-8.

Gray, Peter. 1979. Low attendance slows KHS. Kansas Herpetological Society Newsletter (32):1.

Clarke, Robert F. 1980. Herptiles and fishes of the western Arkansas River in Kansas. United States Army Corps of Engineers, Albuquerque, New Mexico. 55 pp.

Spencer, Dwight. 1980. Spencer, D. 1980. Ross Natural History Reservation: the first twenty years, 1959 to 1979. Emporia State University, Emporia, Kansas.. 64 pp.

Collins, Joseph T. 1980. New records of fishes, amphibians, and reptiles in Kansas for 1979. Technical Publication of the State Biological Survey of Kansas 9:1-11.

Collins, Joseph T. 1981. New records of fishes, amphibians, and reptiles in Kansas for 1980. Technical Publication of the State Biological Survey of Kansas 10:7-19.

Collins, Joseph T. 1982. Report to the Kansas Fish and Game Commission on the status of three amphibians in southeastern Kansas. Kansas Fish and Game Commission, Pratt. 57 pp.

Fitch, Henry S. 1982. Resources of a snake community in prairie-woodland habitat of northeastern Kansas. Pages 83-97 in Herpetological communities: A symposium of the Society for the Study of Amphibians and Reptiles and the Herpetologists League, August 1977.  Wildlife Research Reports 12. 239 pp. U. S. Fish and Wildlife Service, Washington, D. C.

Collins, Joseph T. 1982. Amphibians and Reptiles in Kansas. 2nd edition. University of Kansas Museum of Natural History Public Education Series (8).

Pilch, J. A. 1982. Notes on a brood of Northern Water Snakes from Kansas. Kansas Herpetological Society Newsletter (47):22.

Rogers, Karel L. 1982. Herpetofaunas of the Courland Canal and Hall Ash Local Faunas (Pleistoncene: Early Kansas) of Jewell Co., Kansas. Journal of Herpetology 16(2):174-177.

Miller, Larry L. 1983. Bourbon County field trip well attended and successful. Kansas Herpetological Society Newsletter (54):6-7.

Collins, Joseph T. 1983. New records of fishes, amphibians, and reptiles in Kansas for 1982 . Technical Publication of the State Biological Survey of Kansas 13:9-21.

Achen, Von, Pennie. H., and John L. Rakestraw. 1984. The role of chemoreception in the prey selection of neonate reptiles. Pages 163-172 in Vertebrate Ecology and Systematics— A Tribute to Henry S Fitch. Special Publication Number 10. The University of Kansas Museum of Natural History, Lawrence.

Holman, J. Alan. 1984. Herpetofaunas of the Duck Creek and Williams Local Faunas (Pleistocene: Illinoian) of Kansas. Pages 20-38 in Contributions in Quaternary Vertebrate Paleontology: A Volume in Memorial to John E. Guilday. Special Publication Number 8. Carnegie Museum of Natural History, Pittsburgh, Pennsylvania.

Eshelman, Ralph and Michael Hager. 1984. Two Irvingtonian (Medial Pleistocene) vertebrate faunas from northcentral Kansas. Pages 384-404 in Contributions in Quaternary Vertebrate Paleontology: A Volume in Memorial to John E. Guilday. Special Publication Number 8. Special Publication Number 8, Carnegie Museum of Natural History, Pittsburgh, Pennsylvania.

Brown, Kenneth L. 1984. Pomona: A plains village variant in eastern Kansas and western Missouri. Dissertation. University of Kansas, Lawrence, Kansas. 519 pp.

Heinrich, Mark L. 1984. Herpetofauna of the Konza Prairie Research Natural Area in the Flint Hills region of Kansas with respect to habitat selection. Thesis. Kansas State University, Manhattan, Kansas. 57 pp.

Secor, Stephen M. and Charles C. Carpenter. 1984. Distribution maps of Oklahoma reptiles. Oklahoma Herpetological Society Special Publication (3):1-57.

Capron, Marty B. 1984. Water snakes, lost gold and revolutionaries. Kansas Herpetological Society Newsletter (56):11-14.

Collins, Joseph T. 1984. New records of fishes, amphibians, and reptiles in Kansas for 1983. Kansas Herpetological Society Newsletter (56):15-26.

Lynch, John D. 1985. Annotated checklist of the amphibians and reptiles of Nebraska. Transactions of the Nebraska Academy of Sciences 13:33-57.

Terry, P. A. 1986. Biological survey of the KS segments of Spring River and Shoal Creek. Part 1. Field Survey. Draft. Kansas Fish and Game, Pratt, Kansas. 67 pp.

Collins, Joseph T. 1986. New records of amphibians and reptiles in Kansas for 1986. Kansas Herpetological Society Newsletter (66):9-16.

Holman, J. Alan. 1986. Butler Spring herpetofauna of Kansas (Pleistocene: Illinoian) and its climatic significance. Journal of Herpetology 20(4):568-569.

Coleman, Keith. 1987. Annual KHS Field Trip held at Atchison State Lake. Kansas Herpetological Society Newsletter (68):5-6.

Holman, J. Alan. 1987. Climatic significance of a late Illinoian herpetofauna from southwestern Kansas. Contributions from the Museum of Paleontology, University of Michigan 27(5):129-141.

Holman, J. Alan. 1987. Snakes from the Robert Local Fauna (Late Wisconsinan) of Meade County, Kansas. Contributions from the Museum of Paleontology, University of Michigan 27(6):143-150.

Collins, Joseph T. 1988. New records of amphibians and reptiles in Kansas for 1987. Kansas Herpetological Society Newsletter (71):13-19.

Nulton, Michael T. and Michael S. Rush. 1988. New county records of amphibians and reptiles in Gray County, Kansas. Kansas Herpetological Society Newsletter (74):10-12.

Collins, Joseph T. 1989. New records of amphibians and reptiles in Kansas for 1988. Kansas Herpetological Society Newsletter (75):15-18.

Collins, Joseph T. 1989. First Kansas herp counts held in 1989. Kansas Herpetological Society Newsletter (77):11-.

Collins, Joseph T. 1990. Results of second Kansas herp count held during April-May 1990. Kansas Herpetological Society Newsletter (81):10-12.

Collins, Joseph T. 1990. Maximum size records for Kansas amphibians and reptiles. Kansas Herpetological Society Newsletter (81):13-17.

Conant, Roger and Joseph T. Collins. 1991. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed. Houghton Mifflin Company, Boston, Massachusetts.

Fitch, Henry S. 1991. Reptiles and amphibians of the Kansas ecological reserves. Pages 71-74 in Ecology and Hydrology of Kansas Ecological Reserves and the Baker Wetlands. Multidisciplinary Guidebook 4. Kansas Academy of Science, Lawrence, Kansas.

Collins, Joseph T. 1991. New records of amphibians and reptiles in Kansas for 1990. Kansas Herpetological Society Newsletter (83):7-13.

Capron, Marty B. 1991. Unusual foraging behavior in water snakes (Nerodia) around drying pools in southcentral Kansas. Kansas Herpetological Society Newsletter (84):14-15.

Collins, Joseph T. 1991. Results of third Kansas herp count held during April-May 1991. Kansas Herpetological Society Newsletter (85):9-13.

Collins, Joseph T. 1992. New records of amphibians and reptiles in Kansas for 1991. Kansas Herpetological Society Newsletter (87):12-17.

Taggart, Travis W. 1992. Observations on Kansas amphibians and reptiles. Kansas Herpetological Society Newsletter (88):13-15.

Collins, Joseph T. 1992. Results of the fourth Kansas herp count held during April-May 1992. Kansas Herpetological Society Newsletter (89):10-.

Taggart, Travis W. 1992. Results of the KHS annual field trip to Sheridan County State Lake. Kansas Herpetological Society Newsletter (90):3-4.

Taggart, Travis W. 1992. Nerodia sipedon. Geographic distribution. Herpetological Review 23:91.

Collins, Joseph T. and Suzanne L. Collins. 1993. Amphibians and Reptiles in Kansas. Third Edition. University Press of Kansas, Lawrence, Lawrence. 397 pp.

Collins, Joseph T. and Rundquist, Eric M. 1993. Results of the fifth Kansas herp count held during April-June 1993 . Kansas Herpetological Society Newsletter (94):7-11.

Viets, Brian E. 1993. An annotated list of the herpetofauna of the F. B., and Rena G. Ross Natural History Reservation. Transactions of the Kansas Academy of Science 96(1/2):103-113.

Fitch, Henry S. 1993. Relative abundance of snakes in Kansas. Transactions of the Kansas Academy of Science 96(3/4):213-224.

Rundquist, Eric M. 1994. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14.

Riedle, J. Daren. 1994. A survey of reptiles and amphibians at Montgomery County State Fishing Lake. Kansas Herpetological Society Newsletter (98):11-13.

Rundquist, Eric M. 1994. Additions and corrections [to the results of the sixth annual KHS herp counts held 1 April-31 May 1994]. Kansas Herpetological Society Newsletter (98):4.

Dloogatch, Michael A. (Editor). 1994. Herpetology 1994. Bulletin of the Chicago Herpetological Society 29(12):282-283.

Holman, J. Alan. 1995. Pleistocene Amphibians and Reptiles. Oxford University Press, New York. 243 pp.

Anderson, Lewis, Mark Shaw, Jeff Blodig, and Tom Walker. 1995. Report to the Kansas Department of Wildlife and Parks: Herps encountered during REmap project, summer 1994. Kansas Herpetological Society Newsletter (99):10-17.

Rundquist, Eric M. 1995. Results of the seventh annual KHS herp counts held 1 April-31 May 1995. Kansas Herpetological Society Newsletter (101):11-17.

Rundquist, Eric M. 1995. Additional KHS herp counts for 1995. Kansas Herpetological Society Newsletter (102):11-.

Rundquist, Eric M. 1996. Results of the eighth annual KHS herp counts Held 1 April-31 May 1996. Kansas Herpetological Society Newsletter (104):6-17.

Miller, Larry L. 1996. Third graders conduct amphibian and reptile field study. Kansas Herpetological Society Newsletter (106):15.

Rakestraw, J. 1996. Spring herp counts: A Kansas tradition. Reptile & Amphibian Magazine (March-April):75-80.

Shoup, J. Mark. 1996. Wise as serpents. Kansas Wildlife and Parks 53(4):39.

Rundquist, Eric M. 1997. Results of the ninth annual KHS herp counts held 1 April-31 May 1997. Kansas Herpetological Society Newsletter (108):12-17.

Collins, Joseph T. 1997. A report on the KHS fall field trip to the Marais des Cygnes wildlife refuges. Kansas Herpetological Society Newsletter (110):2-3.

Conant, Roger and Joseph T. Collins. 1998. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed, expanded. Houghton Mifflin Company, Boston, Massachusetts.

Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. 1998. A Key to Amphibians & Reptiles of the Continental United States and Canada. Univ Press of Kansas, Lawrence, Kansas. 131 pp.

Gamble, Jerre. 1998. Marais des Cygnes National Wildlife Refuge Comprehensive Conservation Plan. U.S. Fish and Wildlife Service, Hartford, Kansas. 91 pp.

Riedle, J. Darren. 1998. Water snake feeding records. Kansas Herpetological Society Newsletter (111):16.

Rundquist, Eric M. 1998. Results of the tenth annual KHS herp counts for 1998, held 1 April-31 May. Kansas Herpetological Society Newsletter (112):11-18.

Collins, Joseph T. 1998. Results of the KHS silver anniversary fall field trip. Kansas Herpetological Society Newsletter (114):6-.

Rundquist, Eric M. 1999. Kansas Herpetological Society herp counts: A 10 year summary and evaluation. Kansas Herpetological Society Newsletter (115):42962.

Collins, Joseph T. 1999. New records of amphibians and reptiles in Kansas for 1998. Kansas Herpetological Society Newsletter (116):14-15.

Taggart, Travis W. 1999. Cherokee County fall 1999 herp count. Kansas Herpetological Society Newsletter (117):6.

Fitch, Henry S. 2000. Population structure and biomass of some common snakes in central North America. Scientific Papers of the Natural History Museum, University of Kansas (17):1-7.

Collins, Joseph T. 2000. New records of amphibians and reptiles in Kansas for 1999. Kansas Herpetological Society Newsletter (119):7-9.

Schmidt, Curtis J. 2000. Observations on reptilian predation. Kansas Herpetological Society Newsletter (120):18.

Taggart, Travis W. 2000. KHS spring field trip sets record for attendance. Kansas Herpetological Society Newsletter (120):5-5.

Doren, Van, Mark D., and Curtis J. Schmidt. 2000. A herpetological survey of the Fort Larned National Historic Site, Pawnee County, Kansas. Kansas Herpetological Society Newsletter (120):8-11.

Taggart, Travis W. 2000. Biogeographic analysis of the reptiles (Squamata) in Ellis County, Kansas. Kansas Herpetological Society Newsletter (121):7-16.

Rundquist, Eric M. 2000. Results of the eleventh and twelfth annual KHS herpetofaunal counts for 1999-2000, held 1 April-31 May. Kansas Herpetological Society Newsletter (122):11-16.

Taggart, Travis W. 2000. Results of the KHS 2000 fall field trip. Kansas Herpetological Society Newsletter (122):6-8.

Johnson, Richard W. 2001. Spatial ecology of the Eastern Coachwhip (Masticophis flagellum flagellum) in and eastern Texas upland community. Thesis. Stephen F. Austin State University, Nacogdoches, Texas. 54 pp.

Taggart, Travis W. 2001. The KHS 2001 spring field trip: A rainy rendezvous. Kansas Herpetological Society Newsletter (124):12-14.

Rundquist, Eric M. 2001. Results of the thirteenth annual KHS herp counts for 2001, held 1 April-30 June. Kansas Herpetological Society Newsletter (125):13-16.

Himes, John G. 2002. The role of the Midland Water Snake, Nerodia sipedon, as a competitor, predator, and prey in freshwater communities. Dissertation. University of Southern Mississippi, Hattiesburg, Mississippi. 101 pp.

Kingsbury, Bruce and Joanna Gibson. 2002. Habitat Management Guidelines for Amphibians and Reptiles of the Midwest. Publication of Partners in Amphibian and Reptile Conservation, Address not given. 152 pp.

Fogell, Daniel D. 2002. Occurrence and relative abundance of amphibians and reptiles at Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. Interim Report. National Park Service, Washington, D.C.. 6 pp.

Riedle, J. Daren and A. Hynek. 2002. Amphibian and reptile inventory of the Kansas Army Ammunition Plant, Labette County, Kansas. Journal of Kansas Herpetology (2):18-20.

Ellis, Mark R. 2002. Fall 2002 KHS field trip to Washington County. Journal of Kansas Herpetology (2):4-5.

Taggart, Travis W. 2002. Results of the KHS 2002 fall field Trip. Journal of Kansas Herpetology (4):11-13.

Miller, Larry L. 2002. Shawnee County herp count. Journal of Kansas Herpetology (4):15.

Freeman, Craig C. 2003. A natural areas inventory of the Ft. Leavenworth Military Reservation, Leavenworth County, Kansas. II. Open-file Report No. 117. Kansas Biological Survey, Lawrence, Kansas. 199 pp.

Fogell, Daniel D. 2003. A herpetofaunal inventory of Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. National Park Service, Washington, D.C.. 59 pp.

Suleiman, G. 2003. Fort Riley herpetofaunal count. Journal of Kansas Herpetology (5):11-12.

Platt, Dwight R. 2003. Lizards and snakes (Order Squamata) of Harvey County, Kansas. Journal of Kansas Herpetology (6):13-20.

Fitch, Henry S. 2003. Reproduction in snakes of the Fitch Natural History Reservation in northeastern Kansas. Journal of Kansas Herpetology (6):21-24.

Taggart, Travis W. 2003. Results of the 2003 KHS spring field trip to Wilson County. Journal of Kansas Herpetology (6):2-5.

Miller, Larry L. 2003. Sumner County herp count. Journal of Kansas Herpetology (7):10.

Zerwekh, Mike. 2003. Large Northern Water Snake (Nerodia sipedon) from Kansas. Journal of Kansas Herpetology (7):12.

Suleiman, Gibran. 2003. Fort Riley herp count. Journal of Kansas Herpetology (7):9.

Taggart, Travis W. 2003. Results of the KHS 2003 fall field trip. Journal of Kansas Herpetology (8):14-15.

Fitch, Henry S., Scott Sharp, and Kylee Sharp. 2003. Snakes of the University of Kansas biotic succession area. Journal of Kansas Herpetology (8):20-21.

Gibbons, J. Whitfield and Michael E. Dorcas. 2004. North American Water Snakes. University of Oklahoma Press, Norman.

Delisle, Jennifer M. and William H. Busby. 2004. Biological inventory for vertebrates at Fort Larned National Historic Site of the southern plains network. Natural Heritage Inventory, Kansas Biological Survey, Lawrence. 61 pp.

LaForce, Russell W. 2004. Life history notes: Diet: Nerodia sipedon. Journal of Kansas Herpetology (10):11.

Daniel, James K. 2004. Cherokee County herp count. Journal of Kansas Herpetology (11):10.

Volkmann, Al. 2004. Cowley County herp count. Journal of Kansas Herpetology (11):10.

Gubanyi, James E. 2004. Wilson County herp count. Journal of Kansas Herpetology (11):12.

Taggart, Travis W. 2005. Results of the KHS 2005 fall field trip [to Crawford County]. Journal of Kansas Herpetology (16):19-21.

Taggart, Travis W. 2006. Distribution and status of Kansas herpetofauna in need of information. State Wildlife Grant T7. Kansas Department of Wildlife and Parks, Pratt. vii + 106 pp.

Wilgers, Dustin J., Eva A. Horne, Brett K. Sandercock, and Allan W. Volkmann. 2006. Effects of rangeland management on community dynamics of the herpetofauna of the tallgrass prairie. 62:378-388.

Murrow, Daniel G. 2009. KHS 2009 spring field trip. Journal of Kansas Herpetology (29):42769.

Schmidt, Curtis J. 2009. Geographic distribution: Nerodia sipedon. Journal of Kansas Herpetology (30):11.

Pattishall, Abigail and David Cundall. 2009. Habitat use by synurbic watersnakes (Nerodia sipedon). Herpetologica 65(2):183-198.

Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. 2010. Amphibians, Reptiles, and Turtles of Kansas. Eagle Mountain Publishing., Provo, Utah. 400 pp.

Murrow, Daniel G. 2010. Kansas Herpetological Society spring field trip. Journal of Kansas Herpetology (33):2-3.

Taggart, Travis W. 2011. Kansas Herpetological Society 2011 spring field trip to be held in Chautauqua County. Journal of Kansas Herpetology (37):5-7.

Taggart, Travis W. 2011. Results of the KHS Spring Field Trip to Chautauqua County. Journal of Kansas Herpetology (38):2-4.

Houck, Mike. 2011. Fort Riley Herpetofaunal Survey for 2011. Journal of Kansas Herpetology (39):9.

Rohweder, Megan R. 2012. Spatial conservation prioritization of Kansas for terrestrial vertebrates. Thesis. Fort Hays State University, Hays, Kansas. 151 pp.

Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. 2012. Key to the Herpetofauna of the Continental United States and Canada: Second Edition, Revised and Updated. Univ Press of Kansas, Lawrence, Kansas. 152 pp.

Walley, Harlan D., R. B. King, J. M. Ray, and T. L. Wusterbarth. 2012. Nerodia sipedon. Catalogue of American Amphibians and Reptiles (899):1-58.

Taggart, Travis W. 2013. KHS 2012 Spring Field Trip to Bourbon County State Lake. Collinsorum 2(3/4):3.

Taggart, Travis W. 2013. KHS 2012 Fall Field Trip to Atchison County State Lake. Collinsorum 2(3/4):4.

Taggart, Travis W. 2013. KHS 2013 Fall Field Trip to Butler County State Lake. Collinsorum 2(3/4):6.

Mardis, Dexter and Kevin Scott. 2013. 2013 Kansas Herpetofaunal Counts. Collinsorum 2(3/4):7.

Taggart, Travis W. 2014. Results of the 2014 KHS Fall Field Trip to Woodson County. Collinsorum 3(2-4):12.

Taggart, Travis W. 2014. Recent scientific and standard English name changes effecting the Kansas herpetofauna. Collinsorum 3(2-4):9-10.

Houck, Mike. 2015. 2015 Fort Riley Herpetofaunal Count final report. Collinsorum 4(1):10-11.

Taggart, Travis W. 2015. Spring Field Trip to the Greenhorn Limestone of Russell County. Collinsorum 4(3):2.

Taggart, Travis W. 2015. Summer Field Trip In The Harvey County Sandhills. Collinsorum 4(3):3.

Taggart, Travis W. 2015. Fall Field Trip Held In Washington County. Collinsorum 4(3):4.

Pittman, Galen L., Henry S. Fitch, and W. Dean Kettle. 2016. Vertebrate animals on the Fitch Natural History Reservation (1948-2002). Kansas Biological Survey Report Number 188, Lawrence. 48 pp.

Powell, Robert, Roger Conant, and Joseph T. Collins. 2016. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. Houghton Mifflin Harcourt, Boston. 494 pp.

Taggart, Travis W. 2016. Results of the KHS Summer field trip to Caney River, Chautauqua County, Kansas. Collinsorum 5(2-3):4-5.

Huggins, Donald, Jude Kastens, Debbie Baker, and Craig Freeman. 2017. Conversion of existing farm ponds to wetlands in agricultural landscapes for mitigation, land use treatment and conservation with a perspective toward climate change. Kansas Biological Survey Report No. 189. Kansas Biological Survey, Lawrence. 91 pp.

Crother, Brian I. (editor). 2017. Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, with Comments Regarding Confidence in Our Understanding. Eighth edition. Herpetological Circulars (43):1-102.

Taggart, Travis W. 2017. Results of the 2017 KHS Spring Field Trip to Elk County, Kansas. Collinsorum 6(2-3):6-8.

Houck, Mike. 2018. Herp Count: Fort Riley Military Installation. Collinsorum 7(1):17.

Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. 2019. Key to the Herpetofauna of the Continental United States and Canada. Third Edition. Univ Press of Kansas, Lawrence, Kansas. 192 pp.

Daniel, Richard E. and Brian S. Edmond. 2020. Atlas of Missouri Amphibians and Reptiles for 2019. Privately printed, Columbia, Missouri. 86 pp.

Riedle, J. Daren. 2020. Revisiting Kansas Herpetological Society field trip and Herp Count data: Distributional patterns and trend data of Kansas amphibians and reptiles. Collinsorum 9(1):7-16.

Swartout, Merridith, Philip Vogring, J. Alex Baecher, Chelsea Kross, and John D. Willison. 2020. Prey size and feeding rate do not influence trophic morphology of juvenile water snakes (Nerodia sipedon). Herpetologica 76(1):53–60.

Hullinger, Allison, Zackary Cordes, Daren Riedle, and William Stark. 2020. Habitat assessment of the Broad-headed Skink (Plestiodon laticeps) and the associated squamate community in eastern Kansas. Transactions of the Kansas Academy of Science 123(1-2):137-150.

Taggart, Travis W and Sarah L Taggart. 2021. Herp Count: Neosho County: KHS-2020-01. Collinsorum 9(3):11.

Taggart, Meg, Amelia Jaeger, Jesse J. Taggart, and Travis W. Taggart. 2021. Herp Count: Russell County: KHS-2020-22. Collinsorum 9(3):14-15.

Tyson, Kelsea, Lexis Mader, Thomas Zapletal, Jeremiah Cline, Alyssa Farney, Loegan Hill, Jainee Cowen, Camron Matteson, and David Penning. 2021. Measuring herpetofaunal biodiversity in southwest Missouri. Collinsorum 10(2):13-18.

Nuñez, Leroy P., Levi N Gray, David Weisrock, and Frank T Burbrink. 2023. The phylogenomic and biogeographic history of the Gartersnakes, Watersnakes, and Allies (Natricidae: Thamnophiini). Molecular Phylogenetics and Evolution 186(2023):107844.

Last Updated: 02/19/2026 8:40:43 AM CT

THE CENTER FOR NORTH AMERICAN HERPETOLOGY — Accessed: Tuesday 09 June 2026 13:07 CT