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Kansas Herpetofaunal Atlas

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Ring-necked Snake

Diadophis punctatus (Linnaeus, 1766)

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Overview
HARMLESSLY VENOMOUS. Uses its venom to subdue prey, but is not dangerous to humans because a) they have an ineffective venom delivery mechanism, b) their mouths too small to gain purchase, and/or c) their venom is not adapted for causing physiological damage to mammals.
The Ring-necked Snake is characterized by smooth scales, a divided anal scale, and a bright yellow or orange-yellow ring around the neck. The head, body, and tail are shiny gray-black or blue-black except for the conspicuous neck ring. The belly is yellow, with numerous scattered black dots. The underside of the tail is bright red. Adult males have longer tails than females and have keels or ridges on the scale rows above the cloaca. Females grow larger than males.
Adults normally grow 25.4- 38.0 cm (10-15 inches) in total length. The largest specimen from Kansas is a female (KU 216516) from Shawnee County with a total length of 45.6 cm (17­7⁄8 inches) collected by Mark Ellis on 3 July 1990. The maximum length throughout the range is 70.3 cm (27­11⁄16 inches) (Powell et al., 2016). The maximum weight for a Kansas specimen is 15.2 grams (slightly over ½ ounce).

Distribution
Known from the eastern third of the state. It makes it into the western third along the Cimarron, Arkansas, and Smoky Hill River drainages.

Reproduction
Fitch (1975) studied this snake in northeastern Kansas, and much of the information on this species in Kansas is based on his observations. This species inhabits the rocky hillsides of open woodlands. It is active from March to November, temperatures permitting, but becomes scarce during July and August, probably due to high temperatures and consequent lack of moist surface soil. Ring-necked Snakes apparently prowl at night and spend the day resting on moist soil beneath large rocks. They rarely bask in the sun, preferring to lie beneath sun-warmed rocks to maintain an optimal body temperature. Large numbers of these snakes will congregate beneath a single rock. Why is not known, but it is suspected that this snake uses smell to follow others of its species to the same rock. Henderson (1970) showed that at optimal temperatures (76-85°F) this species requires water about every third day.
During winter, this snake retreats well below the ground surface to avoid cold temperatures. Fitch (1975, 1982) considered this species to be the most abundant vertebrate on his upland study area in Douglas County and obtained estimates of up to 730 Ring-necked Snakes per acre. In his study of 530 Diadophis punctatus in northeast Kansas, Fitch (2000) found that males are larger and live longer than females. He determined that 12 age classes existed in his samples and his density estimates were 1603 individuals per hectare (648/acre) and amounted to 45.9 of the snake biomass in his study area.
Mating occurs during spring, but courtship is unknown. The number of eggs per clutch varies from one to ten (Fitch, 1985), with an average of four; the eggs are laid in late June or early July, probably deep beneath the ground. The eggs hatch in late August or early September. Males become sexually mature during their second season, whereas females require three seasons to reach reproductive maturity. Seigel and Fitch (1985) discovered that this species increased the number of eggs per clutch during a breeding season when there was ample rain the previous autumn, apparently because earthworms, the main diet of this snake in Kansas, were more available for the females to eat.
Henderson (1970) recorded this species feeding primarily on earthworms and small frogs. Fitch (1975) examined the stomach contents of 182 examples of this snake and found 221 earthworms, two insect larvae and one hatchling lizard. He estimated that this snake consumes three times its body weight in earthworms per year. Ring-necked Snakes appear to rely heavily on their sense of smell to locate prey. Fitch (1975) reported predation on the Ring-necked Snake in Douglas County by Copperheads, Timber Rattlesnakes, Racers, Milk Snakes, Bullfrogs, and three species of raptorial birds.
When molested or handled, a Ring-necked Snake may tightly coil and elevate its tail, exposing the bright red undersurface. Smith (1976) demonstrated that large adult males exhibited tail-coiling behavior less often than smaller individuals and females. He felt this behavior provided a defense mechanism for slower-moving juveniles and gravid females. This behavior is defensive and, when accompanied by a cloaca! discharge, may discourage predators, particularly birds.

Remarks
First reported from Kansas by Kennicott (1859) in his description of Diadophis arnyi from Hyatt, Kansas (Anderson County). This specimen was believed to have been collected around 1856, however no date was recorded. That specimen was from the collection of Northwestern University (transferred to the Chicago Academy of Sciences [CHAS] ca. 1920-1925) and ultimately to the United States National Museum (USNM 1968) (Cochran, 1961).
CHAS currently houses one specimen (CHAS 11663) from Hyatt, Anderson County, but it was collected by Edward H. Taylor in 1929 (no other associated data). The type description lists "Samuel Arny" as the collector, however the correct collector is most likely William F. M. Arny, a resident of Hyatt, and a well known scientific collector at the time. The earliest existing specimen (MCZ 9502) with a definitely recorded year, was collected at "Wallace, Kan" by local farmer George R. Allaman and received on 28 December 1885.
In eastern Kansas, it is not uncommon to find twenty or more Ring-necked Snakes under a single piece of cover (rock, board, etc.) during April and May. This species is occasionally observed crossing roads well away from rocky areas and may ultimately be discovered in all but the extreme northwestern corner of the state.
Fifteen of these snakes were found under one rock in Cowley county, Kansas, on 8 April 1933 by Charles E. Burt (Burt and Hoyle, 1935).
Fitch (1975) found that some individuals on his study site survived to an age of fifteen years.

References

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Kennicott, Robert. 1859. Notes on Coluber calligaster of Say, and a description of new species of serpents in the collection of the North Western University of Evanston, Illinois. Proceedings of the Academy of Natural Sciences of Philadelphia 11:98-100.

Cope, Edward D. 1861 "1860". Catalogue of the Colubridae in the Museum of the Academy of Natural Sciences of Philadelphia, with notes and descriptions of new species. Part 2. Proceedings of the Academy of Natural Sciences of Philadelphia 12:241-266.

Jordan, David Starr. 1876. Manual of the Vertebrates of the Northern United States: Including the District East of the Mississippi River, and North of North Carolina and Tennessee, Exclusive of Marine Species. Jansen, McClurg, and Company, Chicago, Illinois.. 342 pp.

Mozley, Annie E. 1877. List of Kansas snakes in the museum of the Kansas State University. Transactions of the Kansas Academy of Science 6:34-35.

Jordan, David S. 1878. Manual of the Vertebrates of the Northern United States: Including the District East of the Mississippi River, and North of North Carolina and Tennessee, Exclusive of Marine Species. Second Edition, Revised, and Enlarged. Jansen, McClurg & Company, Chicago.

Cragin, Francis W. 1880. A preliminary catalogue of Kansas reptiles and batrachians. Transactions of the Kansas Academy of Science 7:112-123.

Yarrow, Henry C. 1882. Check list of North American Reptilia and Batrachia with catalogue of specimens in U. S. National Museum. Bulletin of the United States National Museum (24):1-249.

Garman, Samuel. 1883. The reptiles and batrachians of North America. . Kentucky Geological Survey. Yeoman Press., Frankfort, Kentucky. 185 pp.

Davis, N. S. Jr. and Frank L. Rice. 1883. Descriptive catalogue of North American batrachia and reptilia, found east of Mississippi River. Illinois Natural History Survey Bulletin 1(5):71.

Garman, Samuel. 1883. The reptiles and batrachians of North America. Memiors of the Museum of Comparative Zoology, Harvard University 8(3):xxxi + 1-185.

Stone, Witmer. 1903. A collection of reptiles and batrachians from Arkansas, Indian Territory and western Texas. Proceedings of the Academy of Natural Sciences of Philadelphia 55:538-542.

Branson, Edwin B. 1904. Snakes of Kansas. University of Kansas Science Bulletin 2(13):353-430.

Ditmars, Raymond L. 1907. The Reptile Book; A comprehensive, Popularised Work on the Structure and Habits of the Turtles, Tortoises, Crocodilians, Lizards and Snakes which Inhabit the United States and Northern Mexico. Doubleday, Pae, and Company, New York. 472 pp.

Hurter, Julius. 1911. Herpetology of Missouri. Transactions of the Academy of Science of St. Louis 20(5):59-274.

Linsdale, Jean M. 1925. Land Vertebrates of a Limited Area in Eastern Kansas. Thesis. University of Kansas, Lawrence, Kansas. 312 pp.

Burt, Charles E. 1927. An annotated list of the amphibians and reptiles of Riley County, Kansas. Occasional Papers of the Museum of Zoology, University of Michigan (189):12.

Linsdale, Jean M. 1927. Amphibians and reptiles of Doniphan County, Kansas. Copeia 1927(164):75-81.

Ortenburger, Arthur I. 1928. The whip snakes and racers: Genera Masticophis and Coluber. Memiors of the University of Michigan Museum (1):1-247.

Taylor, Edward H. 1929. A revised checklist of the snakes of Kansas. University of Kansas Science Bulletin 19(5):53-62.

Gloyd, Howard K. 1932. The herpetological fauna of the Pigeon Lake Region, Miami County, Kansas. Papers of the Michigan Academy of Science, Arts and Letters 15:389-408.

Stejneger, Leonhard H. and Thomas Barbour. 1933. A Checklist of North American Amphibians and Reptiles. 3rd Edition. Harvard University Press, Cambridge, Massachusetts.

Burt, Charles E. 1933. Some distributional and ecological records of Kansas reptiles. Transactions of the Academy of Science of St. Louis 26:186-208.

Brennan, Lawrence A. 1934. A check list of the amphibians and reptiles of Ellis County, Kansas. Transactions of the Kansas Academy of Science 37:189-191.

Burt, Charles E. and W. L. Hoyle. 1934. Additional records of the reptiles of the central prairie region of the United States. Transactions of the Kansas Academy of Science 37:193-216.

Brennan, Lawrence A. 1935. Notes on the distribution of Amphibia and Reptilia of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 114 pp.

Burt, Charles E. 1935. Further records of the ecology and distribution of amphibians and reptiles in the middle west. The American Midland Naturalist 16(3):311-366.

Hurd, Myron Alec. 1936. The reptiles of Cherokee County, Kansas. Thesis. Pittsburg State University, Pittsburg, Kansas. 103 pp.

Brumwell, Malcolm J. 1936. Distributional records of the reptilia and amphibians of Kansas. Privately printed. 22 pp.

Brennan, Lawrence A. 1937. A study of the habitat of reptiles and amphibians of Ellis County, Kansas. Transactions of the Kansas Academy of Science 40:341-347.

Schmidt, Karl Peterson and D. D. Davis. 1941. Field Book of Snakes of the United States and Canada. C.P. Putnam and Sons, New York. 365 pp.

Blanchard, Frank N. 1942. The ring-neck snakes, genus Diadophis.. Bulletin of the Chicago Academy of Sciences 7(1):1-144.

Hudson, G. E. 1942. The amphibians and reptiles of Nebraska. Nebraska Conservation Bulletin 24:1-146.

Hall, Henry H. and Hobart M. Smith. 1947. Selected records of reptiles and amphibians from southeastern Kansas. Transactions of the Kansas Academy of Science 49(4):447-454.

Smith, Hobart M. 1950. Handbook of Amphibians and Reptiles of Kansas. University of Kansas, Museum of Natural History, Miscellaneous Publication (2):336.

Brumwell, Malcolm J. 1951. An ecological survey of the Fort Leavenworth Military Reservation. The American Midland Naturalist 45(1):187-231.

Schmidt, Karl P. 1953. A Check List of North American Amphibians and Reptiles. 6th Edition. University of Chicago Press, Chicago, Illinois. 280 pp.

Smith, Hobart M. 1956. Handbook of Amphibians and Reptiles of Kansas. Second edition. University of Kansas Museum of Natural History Miscellaneous Publication (9):1-356.

Loomis, Richard B. 1956. The chigger mites of Kansas (Acarina, Trombiculidae). University of Kansas Science Bulletin 37:1195-1443.

Cochran, Doris M. 1961. Type specimens of reptiles and amphibians in the United States National Museum. Bulletin of the United States National Museum (220):1-289.

Gish, Charles D. 1962. The Herpetofauna of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 34 pp.

Choate, Jerry R. 1967. Wildlife in the Wakarusa Watershed of Northeastern Kansas. Kansas Biological Survey, Lawrence. 46 pp.

Brattstrom, Bayard H. 1967. A succession of Pliocene and Pleistocene snake fauna from the High Plains of the United States. Copeia 1967(1):188-202.

Buikema, Arthur L. and Kenneth B. Armitage. 1969. The effect of temperature on the metabolism of the prairie ringneck snake, Diadophis punctatus arnyi Kennicott. Herpetologica 25(3):194-206.

Branson, Branley A., James Triplett, and Robert Hartmann. 1969. A partial biological survey of the Spring River drainage in Kansas, Oklahoma and Missouri. Part II: The fishes. Transactions of the Kansas Academy of Science 72(4):429-472.

Fitch, Henry S. 1970. Reproductive cycles in lizards and snakes. University of Kansas Museum of Natural History Miscellaneous Publication (52):1-247.

Fitch, Henry S. and Robert R. Fleet. 1970. Natural history of the milk snake (Lampropeltis triangulum) in northeastern Kansas. Herpetologica 26(4):387-396.

Henderson, Robert W. 1974. Resource partitioning among snakes of the University of Kansas Natural History Reservation: A preliminary analysis. Milwaukee Public Museum Contributions in Biology and Geology (1):1-11.

Collins, Joseph T. 1974. Amphibians and Reptiles in Kansas. University of Kansas Museum of Natural History Public Education Series (1):283 pp.

Perry, Janice. 1974. KHS members take trip to southwest Kansas. Kansas Herpetological Society Newsletter (3):2-3.

Pisani, George R. 1974. Herpetology in the KU Division of Biological Sciences. Kansas Herpetological Society Newsletter (4):3-4.

Karns, Daryl, Ray E. Ashton, Jr., and Thomas Swearingen. 1974. Illustrated Guide to Amphibians and Reptiles in Kansas: An Identification Manual. University of Kansas Publications Museum of Natural History Public Education Series(2):viii + 18.

Perry, Janice. 1975. A trip to southeastern Kansas. Kansas Herpetological Society Newsletter (7):4.

Fitch, Henry S. 1975. A demographic study of the Ringneck Snake (Diadophis punctatus) in Kansas. University of Kansas Museum of Natural History Miscellaneous Publication (62):53.

Rundquist, Eric M. 1976. Field checklist (of) amphibians and reptiles of Kansas. Kansas Herpetological Society, Lawrence.

Caldwell, Janalee P. and Gregory. Glass. 1976. Vertebrates of the Woodson County State Fishing Lake and Game Management Area. Pages 62-76 in Preliminary inventory of the biota of Woodson County State Fishing Lake and Game Management Area. Report No. 5. State Biological Survey of Kansas, Lawrence.

Miller, Larry L. 1976. KHS visits Elk County. Kansas Herpetological Society Newsletter (15):1-2.

Grow, David. 1977. Clark County visited by the Society. Kansas Herpetological Society Newsletter (19):1-2.

Trott, Gene. 1977. Chikaskia River wildlife study. Kansas Herpetological Society Newsletter (19):2-3.

Perry, Janice. 1977. KHS members achieve goal: Get Cottonmouth. Kansas Herpetological Society Newsletter (21):3-4.

Curl, Richard L. 1978. Final Environmental Statement: Milford Lake Kansas operation and maintenance. US Army Corps of Engineers, Kansas City District. 158 pp.

Fitch, Henry S. and E. Raymond Hall. 1978. A 20year record of succession on reseeded fields of tallgrass prairie on the Rockefeller Experimental Tract. University of Kansas Museum of Natural History Special Publication (4):1-15.

Warner, M. and R. Wencel. 1978. Chikaskia River study held near Caldwell. Kansas Herpetological Society Newsletter (25):15-16.

Skie, Shelley and Martha Bickford. 1978. KHS takes to the field in July at Winfield. Kansas Herpetological Society Newsletter (26):42798.

Capron, Marty B. 1978. Four county collecting raid: A south central Kansas herping saga. Kansas Herpetological Society Newsletter (26):9-12.

Perry, Janice. 1978. KHS successful at Miami County State Lake. Kansas Herpetological Society Newsletter (27):5.

Collins, Joseph T. and Janalee P. Caldwell. 1978. New records of fishes, amphibians, and reptiles in Kansas for 1977. Technical Publication of the State Biological Survey of Kansas 6:70-88.

Carl, Gary. 1978. Notes on worm eating in the Prairie Ringneck Snake, Diadophis punctatus arnyi. Bulletin of the Maryland Herpetological Society 14(2):95-97.

Clarke, Robert F. 1980. Herptiles and fishes of the western Arkansas River in Kansas. United States Army Corps of Engineers, Albuquerque, New Mexico. 55 pp.

Spencer, Dwight. 1980. Spencer, D. 1980. Ross Natural History Reservation: the first twenty years, 1959 to 1979. Emporia State University, Emporia, Kansas.. 64 pp.

Collins, Joseph T. 1980. New records of fishes, amphibians, and reptiles in Kansas for 1979. Technical Publication of the State Biological Survey of Kansas 9:1-11.

Collins, Joseph T. 1981. New records of fishes, amphibians, and reptiles in Kansas for 1980. Technical Publication of the State Biological Survey of Kansas 10:7-19.

Fitch, Henry S. 1981. Sexual size differences in reptiles. University of Kansas Museum of Natural History Miscellaneous Publication 70:71.

Collins, Joseph T. 1982. Report to the Kansas Fish and Game Commission on the status of three amphibians in southeastern Kansas. Kansas Fish and Game Commission, Pratt. 57 pp.

Fitch, Henry S. 1982. Resources of a snake community in prairie-woodland habitat of northeastern Kansas. Pages 83-97 in Herpetological communities: A symposium of the Society for the Study of Amphibians and Reptiles and the Herpetologists League, August 1977.  Wildlife Research Reports 12. 239 pp. U. S. Fish and Wildlife Service, Washington, D. C.

Collins, Joseph T. 1982. Amphibians and Reptiles in Kansas. 2nd edition. University of Kansas Museum of Natural History Public Education Series (8).

Brannian, John D. 1983. Metabolic effects of temperature and thyroid activity in the Prairie Ringneck Snake, Diadophis punctatus arnyi. Thesis. University of Nebraska at Omaha, Omaha, Nebraska. 44 pp.

Trott, Gene. 1983. Chikaskia River wildlife study. Kansas Herpetological Society Newsletter (52):3-4.

Fraser, John C. 1983. A trip to the 'TransPecos'. Kansas Herpetological Society Newsletter (54):18-23.

Collins, Joseph T. 1983. New records of fishes, amphibians, and reptiles in Kansas for 1982 . Technical Publication of the State Biological Survey of Kansas 13:9-21.

Achen, Von, Pennie. H., and John L. Rakestraw. 1984. The role of chemoreception in the prey selection of neonate reptiles. Pages 163-172 in Vertebrate Ecology and Systematics— A Tribute to Henry S Fitch. Special Publication Number 10. The University of Kansas Museum of Natural History, Lawrence.

Brown, Kenneth L. 1984. Pomona: A plains village variant in eastern Kansas and western Missouri. Dissertation. University of Kansas, Lawrence, Kansas. 519 pp.

Heinrich, Mark L. 1984. Herpetofauna of the Konza Prairie Research Natural Area in the Flint Hills region of Kansas with respect to habitat selection. Thesis. Kansas State University, Manhattan, Kansas. 57 pp.

Secor, Stephen M. and Charles C. Carpenter. 1984. Distribution maps of Oklahoma reptiles. Oklahoma Herpetological Society Special Publication (3):1-57.

Schwarting, Nancy. 1984. KHS field trip, May 1984. Kansas Herpetological Society Newsletter (57):3-4.

Collins, Joseph T. 1984. New records of fishes, amphibians and reptiles in Kansas for 1984. Kansas Herpetological Society Newsletter (58):14-20.

Miller, Larry L. 1985. KHS 1985 field trip to Kirwin Reservoir. Kansas Herpetological Society Newsletter (61):11-12.

Fitch, Henry S. 1985. Variation in clutch and litter size in New World reptiles. University of Kansas Museum of Natural History Miscellaneous Publication (76):1-76.

Lynch, John D. 1985. Annotated checklist of the amphibians and reptiles of Nebraska. Transactions of the Nebraska Academy of Sciences 13:33-57.

Collins, Joseph T. 1986. New records of amphibians and reptiles in Kansas for 1985. Kansas Herpetological Society Newsletter (63):4.

Collins, Joseph T. 1986. New records of amphibians and reptiles in Kansas for 1986. Kansas Herpetological Society Newsletter (66):9-16.

Holman, J. Alan. 1986. Butler Spring herpetofauna of Kansas (Pleistocene: Illinoian) and its climatic significance. Journal of Herpetology 20(4):568-569.

Coleman, Keith. 1987. Annual KHS Field Trip held at Atchison State Lake. Kansas Herpetological Society Newsletter (68):5-6.

Simmons, John E. 1987. September 1987 field trip report. Kansas Herpetological Society Newsletter (69):42894.

Holman, J. Alan. 1987. Climatic significance of a late Illinoian herpetofauna from southwestern Kansas. Contributions from the Museum of Paleontology, University of Michigan 27(5):129-141.

Holman, J. Alan. 1987. Snakes from the Robert Local Fauna (Late Wisconsinan) of Meade County, Kansas. Contributions from the Museum of Paleontology, University of Michigan 27(6):143-150.

Miller, Larry L. 1988. Harper County KHS field trip well attended. Kansas Herpetological Society Newsletter (72):5-6.

Collins, Joseph T. 1989. New records of amphibians and reptiles in Kansas for 1988. Kansas Herpetological Society Newsletter (75):15-18.

Capron, Marty B. 1989. Threatened and endangered: A critique of the Kansas list. Kansas Herpetological Society Newsletter (76):14-15.

Collins, Joseph T. 1989. First Kansas herp counts held in 1989. Kansas Herpetological Society Newsletter (77):11-.

Collins, Joseph T. 1989. New records of amphibians and reptiles in Kansas for 1989. Kansas Herpetological Society Newsletter (78):16-21.

Collins, Joseph T. 1990. Results of second Kansas herp count held during April-May 1990. Kansas Herpetological Society Newsletter (81):10-12.

Collins, Joseph T. 1990. Maximum size records for Kansas amphibians and reptiles. Kansas Herpetological Society Newsletter (81):13-17.

Joy, Jack. 1990. An additional note on Howard K. Gloyd. Bulletin of the Chicago Herpetological Society 25(10):180.

Conant, Roger and Joseph T. Collins. 1991. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed. Houghton Mifflin Company, Boston, Massachusetts.

Fitch, Henry S. 1991. Reptiles and amphibians of the Kansas ecological reserves. Pages 71-74 in Ecology and Hydrology of Kansas Ecological Reserves and the Baker Wetlands. Multidisciplinary Guidebook 4. Kansas Academy of Science, Lawrence, Kansas.

Collins, Joseph T. 1991. New records of amphibians and reptiles in Kansas for 1990. Kansas Herpetological Society Newsletter (83):7-13.

Collins, Joseph T. 1991. Results of third Kansas herp count held during April-May 1991. Kansas Herpetological Society Newsletter (85):9-13.

Collins, Joseph T. 1992. Results of the fourth Kansas herp count held during April-May 1992. Kansas Herpetological Society Newsletter (89):10-.

Taggart, Travis W. 1992. Results of the KHS annual field trip to Sheridan County State Lake. Kansas Herpetological Society Newsletter (90):3-4.

Rundquist, Eric M. 1992. Results of the KHS 1992 fall field trip. Kansas Herpetological Society Newsletter (90):4.

Young, Eugene A. 1993. A Survey of the Vertebrates of Slate Creek Salt Marsh, Sumner County, Kansas, with an Emphasis on Waterbirds. Thesis. Fort Hays State University, Hays, Kansas. 189 pp.

Collins, Joseph T. and Suzanne L. Collins. 1993. Amphibians and Reptiles in Kansas. Third Edition. University Press of Kansas, Lawrence, Lawrence. 397 pp.

Collins, Joseph T. and Rundquist, Eric M. 1993. Results of the fifth Kansas herp count held during April-June 1993 . Kansas Herpetological Society Newsletter (94):7-11.

Viets, Brian E. 1993. An annotated list of the herpetofauna of the F. B., and Rena G. Ross Natural History Reservation. Transactions of the Kansas Academy of Science 96(1/2):103-113.

Fitch, Henry S. 1993. Relative abundance of snakes in Kansas. Transactions of the Kansas Academy of Science 96(3/4):213-224.

Fitzgerald, Eve C. and Charles Nilon. 1994. Classification of habitats for endangered and threatened species in Wyandotte County, Kansas. Kansas Department of Wildlife and Parks, Pratt, Kansas. 98 pp.

Collins, Joseph T. 1994. New records of amphibians and reptiles in Kansas for 1993. Kansas Herpetological Society Newsletter (97):15-19.

Rundquist, Eric M. 1994. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14.

Riedle, J. Daren. 1994. A survey of reptiles and amphibians at Montgomery County State Fishing Lake. Kansas Herpetological Society Newsletter (98):11-13.

Rundquist, Eric M. 1994. Additions and corrections [to the results of the sixth annual KHS herp counts held 1 April-31 May 1994]. Kansas Herpetological Society Newsletter (98):4.

Dloogatch, Michael A. (Editor). 1994. Herpetology 1994. Bulletin of the Chicago Herpetological Society 29(12):282-283.

Holman, J. Alan. 1995. Pleistocene Amphibians and Reptiles. Oxford University Press, New York. 243 pp.

Rundquist, Eric M. 1995. Results of the seventh annual KHS herp counts held 1 April-31 May 1995. Kansas Herpetological Society Newsletter (101):11-17.

Rundquist, Eric M. 1995. Additional KHS herp counts for 1995. Kansas Herpetological Society Newsletter (102):11-.

Miller, Larry L. 1996. Results of the KHS 1995 fall field trip. Kansas Herpetological Society Newsletter (103):3.

Rundquist, Eric M. 1996. Results of the eighth annual KHS herp counts Held 1 April-31 May 1996. Kansas Herpetological Society Newsletter (104):6-17.

Miller, Larry L. 1996. Third graders conduct amphibian and reptile field study. Kansas Herpetological Society Newsletter (106):15.

Miller, Larry L. 1996. Many amphibian and reptile species identified during KHS 1996 fall field trip to Wabaunsee County. Kansas Herpetological Society Newsletter (106):2-3.

Rakestraw, J. 1996. Spring herp counts: A Kansas tradition. Reptile & Amphibian Magazine (March-April):75-80.

Rundquist, Eric M. 1997. Results of the ninth annual KHS herp counts held 1 April-31 May 1997. Kansas Herpetological Society Newsletter (108):12-17.

Rundquist, Eric M. 1997. Addendum to 1997 KHS herp counts. Kansas Herpetological Society Newsletter (109):14-15.

Collins, Joseph T. 1997. A report on the KHS fall field trip to the Marais des Cygnes wildlife refuges. Kansas Herpetological Society Newsletter (110):2-3.

Clark, Donald R., Jr., Christine M. Bunck, and Russell J. Hall. 1997. Female reproductive dynamics in a Maryland population of ringneck snakes (Diadophis punctatus). Journal of Herpetology 31(4):476-483.

Conant, Roger and Joseph T. Collins. 1998. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed, expanded. Houghton Mifflin Company, Boston, Massachusetts.

Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. 1998. A Key to Amphibians & Reptiles of the Continental United States and Canada. Univ Press of Kansas, Lawrence, Kansas. 131 pp.

Rundquist, Eric M. 1998. Results of the tenth annual KHS herp counts for 1998, held 1 April-31 May. Kansas Herpetological Society Newsletter (112):11-18.

Simmons, John E. 1998. A technique to remove reptiles from glue boards. Kansas Herpetological Society Newsletter (114):18.

Collins, Joseph T. 1998. Results of the KHS silver anniversary fall field trip. Kansas Herpetological Society Newsletter (114):6-.

Fitch, Henry S. 1999. A Kansas Snake Community: Composition and Changes over 50 Years. Krieger Publishing Company, Malabar, Florida.

Rundquist, Eric M. 1999. Kansas Herpetological Society herp counts: A 10 year summary and evaluation. Kansas Herpetological Society Newsletter (115):42962.

Taggart, Travis W. 1999. Cherokee County fall 1999 herp count. Kansas Herpetological Society Newsletter (117):6.

Fitch, Henry S. 2000. Population structure and biomass of some common snakes in central North America. Scientific Papers of the Natural History Museum, University of Kansas (17):1-7.

Taggart, Travis W. 2000. KHS spring field trip sets record for attendance. Kansas Herpetological Society Newsletter (120):5-5.

Taggart, Travis W. 2000. Biogeographic analysis of the reptiles (Squamata) in Ellis County, Kansas. Kansas Herpetological Society Newsletter (121):7-16.

Rundquist, Eric M. 2000. Results of the eleventh and twelfth annual KHS herpetofaunal counts for 1999-2000, held 1 April-31 May. Kansas Herpetological Society Newsletter (122):11-16.

Taggart, Travis W. 2000. Results of the KHS 2000 fall field trip. Kansas Herpetological Society Newsletter (122):6-8.

Guarisco, Hank. 2001. Ode to an ophidion autumn. Kansas Herpetological Society Newsletter (123):19.

Taggart, Travis W. 2001. The KHS 2001 spring field trip: A rainy rendezvous. Kansas Herpetological Society Newsletter (124):12-14.

Rundquist, Eric M. 2001. Results of the thirteenth annual KHS herp counts for 2001, held 1 April-30 June. Kansas Herpetological Society Newsletter (125):13-16.

Kingsbury, Bruce and Joanna Gibson. 2002. Habitat Management Guidelines for Amphibians and Reptiles of the Midwest. Publication of Partners in Amphibian and Reptile Conservation, Address not given. 152 pp.

Fogell, Daniel D. 2002. Occurrence and relative abundance of amphibians and reptiles at Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. Interim Report. National Park Service, Washington, D.C.. 6 pp.

Ellis, Mark R. 2002. Fall 2002 KHS field trip to Washington County. Journal of Kansas Herpetology (2):4-5.

Taggart, Travis W. 2002. Results of the KHS 2002 fall field Trip. Journal of Kansas Herpetology (4):11-13.

Miller, Larry L. 2002. Shawnee County herp count. Journal of Kansas Herpetology (4):15.

Miller, Larry L. 2002. Sumner County herp count. Journal of Kansas Herpetology (4):15.

Rundquist, Eric M. 2002. Natural history of the Night Snake, Hypsiglena torquata, in Kansas. Journal of Kansas Herpetology (4):16-20.

Freeman, Craig C. 2003. A natural areas inventory of the Ft. Leavenworth Military Reservation, Leavenworth County, Kansas. II. Open-file Report No. 117. Kansas Biological Survey, Lawrence, Kansas. 199 pp.

Fogell, Daniel D. 2003. A herpetofaunal inventory of Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. National Park Service, Washington, D.C.. 59 pp.

Suleiman, G. 2003. Fort Riley herpetofaunal count. Journal of Kansas Herpetology (5):11-12.

Taggart, Travis W. 2003. KHS conducts first systematic road survey. Journal of Kansas Herpetology (6):11-12.

Volkmann, Allan W. 2003. Predation by a Ringneck Snake. Journal of Kansas Herpetology (6):12.

Platt, Dwight R. 2003. Lizards and snakes (Order Squamata) of Harvey County, Kansas. Journal of Kansas Herpetology (6):13-20.

Fitch, Henry S. 2003. Reproduction in snakes of the Fitch Natural History Reservation in northeastern Kansas. Journal of Kansas Herpetology (6):21-24.

Taggart, Travis W. 2003. Results of the 2003 KHS spring field trip to Wilson County. Journal of Kansas Herpetology (6):2-5.

Miller, Larry L. 2003. Sumner County herp count. Journal of Kansas Herpetology (7):10.

Miller, Larry L. and Suzanne L. Miller. 2003. Wakarusa herp count. Journal of Kansas Herpetology (7):10.

Volkmann, Al. 2003. Cowley County herp count 1. Journal of Kansas Herpetology (7):7.

Lokke, John L. and Jill Lokke. 2003. Cowley County herp count 2. Journal of Kansas Herpetology (7):8.

Suleiman, Gibran. 2003. Fort Riley herp count. Journal of Kansas Herpetology (7):9.

Taggart, Travis W. 2003. Results of the KHS 2003 fall field trip. Journal of Kansas Herpetology (8):14-15.

Fitch, Henry S., Scott Sharp, and Kylee Sharp. 2003. Snakes of the University of Kansas biotic succession area. Journal of Kansas Herpetology (8):20-21.

Fitch, Henry S. 2004. The effect of female size on number of eggs or young in snakes. Journal of Kansas Herpetology (9):11-12.

Taggart, Travis W. 2004. Kansas Herpetological Society 2004 spring field trip. Journal of Kansas Herpetology (9):2.

McNearney, Ross. 2004. Life history. Diadophis punctatus. Albino. Journal of Kansas Herpetology (10):11.

Fitch, Henry S. 2004. Food surplus and body size in local populations of snakes. Journal of Kansas Herpetology (10):14-16.

Taggart, Travis W. 2004. Results of the 2004 KHS spring field trip to Logan County. Journal of Kansas Herpetology (10):2-7.

Daniel, James K. 2004. Cherokee County herp count. Journal of Kansas Herpetology (11):10.

Volkmann, Al. 2004. Cowley County herp count. Journal of Kansas Herpetology (11):10.

Washburne, M. 2004. Ellsworth County herp count. Journal of Kansas Herpetology (11):10.

Collins, Joseph T. 2004. Marais des Cygnes herp count. Journal of Kansas Herpetology (11):11.

Miller, Larry L. 2004. Sumner County herp count. Journal of Kansas Herpetology (11):11-12.

Ellis, Mark R. and Kathy Ellis. 2004. Wakarusa herp count. Journal of Kansas Herpetology (11):12.

Taggart, Travis W. 2004. Results of the KHS 2004 fall feld trip . Journal of Kansas Herpetology (12):15-16.

Fitch, Henry S. 2004. Observations on Ringneck Snakes (Diadophis punctatus). Journal of Kansas Herpetology (12):19.

Willson, John D. and Michael E. Dorcas. 2004. Aspects of the ecology of small fossorial snakes in the western piedmont of North Carolina. Southeastern Naturalist 3(1):1-12.

Taggart, Travis W. and Curtis J. Schmidt. 2005. Geographic distribution: Diadophis punctatus. Journal of Kansas Herpetology (14):11.

Taggart, Travis W. 2005. Results of the KHS 2005 fall field trip [to Crawford County]. Journal of Kansas Herpetology (16):19-21.

Taggart, Travis W. 2006. Distribution and status of Kansas herpetofauna in need of information. State Wildlife Grant T7. Kansas Department of Wildlife and Parks, Pratt. vii + 106 pp.

Miller, Larry L. 2006. Late fall ringneck snake activity. Journal of Kansas Herpetology (17):6.

Wilgers, Dustin J. and Eva A. Horne. 2006. Effects of different burn regimes on tallgrass prairie herpetofaunal species diversity and community composition in the Flint Hills, Kansas. Journal of Herpetology 40:73-84.

Gray, Brian S. 2006. Artificial hibernation of some temperate North American snakes. Bulletin of the Chicago Herpetological Society 41(6):101-105.

Dloogatch, Michael A. (Editor). 2006. Herpetology 2006. Bulletin of the Chicago Herpetological Society 41(6):111-114.

Wilgers, Dustin J., Eva A. Horne, Brett K. Sandercock, and Allan W. Volkmann. 2006. Effects of rangeland management on community dynamics of the herpetofauna of the tallgrass prairie. 62:378-388.

Taggart, Travis W., Joseph T. Collins, and Curtis J. Schmidt. 2007. Estimates of amphibian, reptile, and turtle mortality if Phostoxin is applied to 10,000 acres of prairie dog burrows in Logan County, Kansas. Kansas Department of Wildlife and Parks, Pratt. 5 pp.

Taggart, Travis W. 2008. KHS 2008 spring field trip. Journal of Kansas Herpetology (25):2-3.

Collins, Joseph T. 2008. Ringneck Snake revelations. Journal of Kansas Herpetology (26):7-8.

Meshaka, Walter E., Jr. 2008. Seasonal activity, reproduction, and growth of the Ringneck Snake (Diadophis punctatus) in Pennsylvania. Journal of Kansas Herpetology 28:17-20.

Fontanella, Frank M., Chris R. Feldman, Mark E. Siddall, and Frank T. Burbrink. 2008. Phylogeography of Diadophis punctatus: Extensive lineage diversity and repeated patterns of historical demography in a transcontinental snake. Molecular Phylogenetics and Evolution 46:1049-1070.

Todd, Brian D., John D. Willson, Christopher T. Winne, Raymond D. Semlitsch, and J. Whitfield Gibbons. 2008. Ecology of the Southeastern Crowned Snake, Tantilla coronata. Copeia 2008(2):388-394.

Nolting, Ray. 2008. Snake hunter finds rattlesnake in Neosho. Parsons Sun 1 May 2008:1,8.

Fontanella, Frank M. 2009. Evolution, phylogeography and species boundaries of the Ringneck Snake Diadophis. Dissertation. City University of New York, New York, New York. 211 pp.

Murrow, Daniel G. 2009. KHS 2009 spring field trip. Journal of Kansas Herpetology (29):42769.

Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. 2010. Amphibians, Reptiles, and Turtles of Kansas. Eagle Mountain Publishing., Provo, Utah. 400 pp.

Murrow, Daniel G. 2010. Kansas Herpetological Society spring field trip. Journal of Kansas Herpetology (33):2-3.

Gray, Brian S. 2010. Distribution of native and exotic earthworms in the eastern United States: Implications for the ecology of vermivorous snakes. Bulletin of the Chicago Herpetological Society 45(5):73-86.

McMartin, David C. 2011. U. S. Army 2011 Fort Leavenworth Herpetofaunal Survey: 23 April - 09 May 2011. Privately printed, Leavenworth, Kansas. 33 pp.

Busby, William. 2011. Ecology of the Smooth Earth Snake (Virginia valeriae) and Redbelly Snake (Storeria occipitomaculata) in Northeastern Kansas. Open-file Report No. 172. Kansas Department of Wildlife and Parks, Pratt, Kansas. 37 pp.

Taggart, Travis W. 2011. Kansas Herpetological Society 2011 spring field trip to be held in Chautauqua County. Journal of Kansas Herpetology (37):5-7.

Taggart, Travis W. 2011. Results of the KHS Spring Field Trip to Chautauqua County. Journal of Kansas Herpetology (38):2-4.

Taggart, Travis W. 2011. Results of the Kansas Herpetological Society 2011 Summer Field Trip to Scott State Park. Journal of Kansas Herpetology (39):2.

Taggart, Travis W. 2011. Results of the Kansas Herpetological Society 2011 Fall Field Trip to Lovewell State Park. Journal of Kansas Herpetology (39):4-5.

McMartin, D. Chris. 2011. Herp Count: Fort Leavenworth Herpetofaunal Survey for 2011. Journal of Kansas Herpetology (39):8-9.

Houck, Mike. 2011. Fort Riley Herpetofaunal Survey for 2011. Journal of Kansas Herpetology (39):9.

Rohweder, Megan R. 2012. Spatial conservation prioritization of Kansas for terrestrial vertebrates. Thesis. Fort Hays State University, Hays, Kansas. 151 pp.

Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. 2012. Key to the Herpetofauna of the Continental United States and Canada: Second Edition, Revised and Updated. Univ Press of Kansas, Lawrence, Kansas. 152 pp.

Hamilton, Bryan Tyler, Rachel Hart, and Jack W. Sites Jr. 2012. Feeding ecology of the Milksnake (Lampropeltis triangulum, Colubridae) in the western United States. Journal of Herpetology 46(4):515-522.

Sinclair, Tom. 2013. A four-day spring snake count across northern Kansas. Collinsorum 2(1/2):9.

Baldwin, Mary Kate. 2013. “Herps in Havensville” Herp Surveys. Collinsorum 2(3/4):10.

Taggart, Travis W. 2013. KHS 2012 Spring Field Trip to Bourbon County State Lake. Collinsorum 2(3/4):3.

Taggart, Travis W. 2013. KHS 2012 Fall Field Trip to Atchison County State Lake. Collinsorum 2(3/4):4.

Taggart, Travis W. 2013. KHS 2013 Spring Field Trip to Schermerhorn Park, Cherokee County. Collinsorum 2(3/4):4.

Taggart, Travis W. 2013. KHS 2013 Fall Field Trip to Butler County State Lake. Collinsorum 2(3/4):6.

Johnson, Stephen R. 2013. Half-time herping on one big prairie. Bulletin of the Chicago Herpetological Society 48(5):65-66.

McMartin, D. Chris. 2014. Fort Leavenworth Heretofaunal Survey for 2013. Collinsorum 3(1):10.

Taggart, Travis W. 2014. Results of the 2014 KHS Spring Field Trip to Barber County. Collinsorum 3(2-4):11.

Taggart, Travis W. 2014. Results of the 2014 KHS Fall Field Trip to Woodson County. Collinsorum 3(2-4):12.

Pauly-Hubbard, Emma C. 2014. Prairie Ring-necked Snake Diadophis punctatus arnyi Captive Care,. Collinsorum 3(2-4):17-18.

Taggart, Travis W. 2014. Recent scientific and standard English name changes effecting the Kansas herpetofauna. Collinsorum 3(2-4):9-10.

Houck, Mike. 2015. 2015 Fort Riley Herpetofaunal Count final report. Collinsorum 4(1):10-11.

Taggart, Travis W. 2015. Spring Field Trip to the Greenhorn Limestone of Russell County. Collinsorum 4(3):2.

Taggart, Travis W. 2015. Fall Field Trip Held In Washington County. Collinsorum 4(3):4.

Pittman, Galen L., Henry S. Fitch, and W. Dean Kettle. 2016. Vertebrate animals on the Fitch Natural History Reservation (1948-2002). Kansas Biological Survey Report Number 188, Lawrence. 48 pp.

Powell, Robert, Roger Conant, and Joseph T. Collins. 2016. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. Houghton Mifflin Harcourt, Boston. 494 pp.

Taggart, Travis W. 2016. Spring 2016 KHS field trip to Clark County was a soggy success. Collinsorum 5(2-3):2-3.

Taggart, Travis W. 2016. Results of the KHS Summer field trip to Caney River, Chautauqua County, Kansas. Collinsorum 5(2-3):4-5.

Crother, Brian I. (editor). 2017. Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, with Comments Regarding Confidence in Our Understanding. Eighth edition. Herpetological Circulars (43):1-102.

Mardis, Dexter R. 2017. Results from three Herpetofaunal tallies at Wichita State University’s Youngmeyer Ranch in Northwestern Elk County. Collinsorum 6(1):8-10.

Taggart, Travis W. 2017. Results of the 2017 KHS Spring Field Trip to Elk County, Kansas. Collinsorum 6(2-3):6-8.

Schmidt, Curtis J. 2017. Herp Count: Southeast Ellis County. Collinsorum 6(2-3):9.

Taggart, Travis W. 2017. Herp Count: Clark County State Lake. Collinsorum 6(2-3):9.

Taggart, Travis W. 2017. Herp Count: Northeast Barton County. Collinsorum 6(2-3):9.

Houck, Mike. 2018. Herp Count: Fort Riley Military Installation. Collinsorum 7(1):17.

Leao, Suelem Muniz, Eric R. Pianka, and Nicolás Pelegrin. 2018. Is there evidence for population regulation in amphibians and reptiles? Journal of Herpetology 52(1):28-33.

Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. 2019. Key to the Herpetofauna of the Continental United States and Canada. Third Edition. Univ Press of Kansas, Lawrence, Kansas. 192 pp.

Degano, Maria E. 2019. The influence of landscape on genetic divergence of the Prairie Ring-necked Snake (Diadophis punctatus arnyi) in Missouri. Thesis. Truman State University, Kirksville, Missouri. 69 pp.

Daniel, Richard E. and Brian S. Edmond. 2020. Atlas of Missouri Amphibians and Reptiles for 2019. Privately printed, Columbia, Missouri. 86 pp.

Riedle, J. Daren. 2020. Revisiting Kansas Herpetological Society field trip and Herp Count data: Distributional patterns and trend data of Kansas amphibians and reptiles. Collinsorum 9(1):7-16.

Hullinger, Allison, Zackary Cordes, Daren Riedle, and William Stark. 2020. Habitat assessment of the Broad-headed Skink (Plestiodon laticeps) and the associated squamate community in eastern Kansas. Transactions of the Kansas Academy of Science 123(1-2):137-150.

Taggart, Travis W and Sarah L Taggart. 2021. Herp Count: Neosho County: KHS-2020-01. Collinsorum 9(3):11.

Taggart, Travis W and Sarah L Taggart. 2021. Herp Count: Cherokee County: KHS-2020-02. Collinsorum 9(3):11-12.

Abbott, Samuel S. 2021. Herp Count: Cowley County: KHS-2020-06. Collinsorum 9(3):12.

Riedle, J. Daren. 2021. Herp Count: Pratt County: KHS-2020-07. Collinsorum 9(3):12.

Schmidt, Curtis J and Avery Schmidt. 2021. Herp Count: KHS-2020-4. Collinsorum 9(3):12.

Mardis, Dexter R. 2021. Herp Count: Sumner County: KHS-2020-08. Collinsorum 9(3):12-13.

Taggart, Travis W and Sarah L Taggart. 2021. Herp Count: Hodgeman County: KHS-2020-09. Collinsorum 9(3):13.

Taggart, Travis W and Sarah L Taggart. 2021. Herp Count: Seward County: KHS-2020-13. Collinsorum 9(3):13.

Schmidt, Curtis J. 2021. Herp Count: Russell County: KHS-2020-19. Collinsorum 9(3):14.

Riedle, J. Daren, Tamera D. Riedle, Zachary Riedle, and Greya Riedle. 2021. Herp Count: Montgomery County: KHS-2020-23. Collinsorum 9(3):14-15.

Taggart, Travis W. 2021. Herp Count: Russell County: KHS-2020-24. Collinsorum 9(3):14-15.

Taggart, Megan M. and Travis W. Taggart. 2021. Herp Count: Seward County: KHS-2020-27. Collinsorum 9(3):15.

Taggart, Travis W., Dan Fogell, and Christopher Visser. 2021. Herp Count: Russell County: KHS-2020-25. Collinsorum 9(3):15.

Underwood, Tyler L. and Keith Geluso. 2022. Geographic distribution: Diadophis punctatus. USA, Nebraska. Herpetological Review 53(1):73-4.

Ward, Krista J. 2022. Geographic distribution: Diadophis punctatus. USA, Kansas. Herpetological Review 53(2):263.

Wright, Jake. 2024. Herpetofaunal assemblage at a former rock quarry in the Flint Hills of Kansas. Collinsorum 13(1):8-9.

Last Updated: 02/25/2024 10:26:43 PM CT

THE CENTER FOR NORTH AMERICAN HERPETOLOGY — Accessed: Tuesday 09 June 2026 13:15 CT