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Kansas Herpetofaunal Atlas

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Western Wormsnake

Carphophis vermis (Kennicott, 1859)

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Overview
HARMLESS. The Western Wormsnake is characterized by smooth scales, a divided anal plate, and a distinctive color. The head, body, and tail are black or gray-black and contrast sharply with the belly color of bright reddish pink, which extends a short distance up onto the sides. Male Western Wormsnakes have longer tails than females and have keels or ridges on the scale rows above the cloaca! opening. Females grow larger than males.
Adults normally grow 19.0-28.0 cm (7½-11 inches) in total length. The largest specimen from Kansas is a female (KU 206169) from Doniphan County with a total length of 36.6 cm (14­3⁄8 inches) collected by Robert Powell on 2 May 1986. The maximum length throughout the range is 39.1 cm (15­3⁄8 inches) (Powell et al., 2016) The maximum weight for Kansas specimen is 14.5 grams (½ ounce).

Distribution
Found in the more heavily forested eastern third of Kansas. This species favors more heavily forested areas and makes it into the Flint Hills along riparian corridors.

Reproduction
Clark (1967, 1970) studied this small, secretive snake in northeastern Kansas, and much of the information from Kansas is based on his observations.
Western Wormsnakes are found beneath limestone rocks in the loose, damp soil of wooded or partly wooded hillsides. Some have been observed at woodland edge. This reptile is active from March to early October, with a peak observable abundance during April. Few specimens are seen during mid to late summer as temperatures increase and the soil under cover become dry. During winter, Western Wormsnakes retreat beneath the soil to depths of at least two feet to avoid cold temperatures.
This species is active day and night at air temperatures ranging from 8.9° to 25.6°C (48° to 78°F), but its daily activities are more dependent on soil temperature. These snakes have home ranges varying from 65.0 to 46.5 square meters (700 to 7,500 square feet), and males have much larger home ranges than females. Population density for this snake in northeastern Kansas is estimated to be 150- 300 individuals per acre within suitable habitat. When burrowing through soil, this snake pushes and rotates its head into areas of least resistance, such as crevices and cracks. This species is rarely discovered out from under cover. Smith (1950, 1956) observed a specimen active on the surface along a creek in Cherokee County. During his two-year study, Taggart (2006) noted 41 specimens under cover and one road-killed specimen.
Western Wormsnakes mate during the spring (April and May) and again in fall (September and October). If courtship exists, it is unknown. Females normally lay an average of one to five eggs per clutch (Fitch, 1985) from mid-June to early July. The eggs, presumably laid in burrows beneath the soil or under rocks, are known to hatch in August.
Dloogatch (1978) reported the discovery of a clutch of five eggs in Douglas County. The eggs were beneath a limestone rock on a south-facing hillside, and all hatched on 13- 14 August.
Clark (1970) and Henderson (1974) reported the Western Wormsnake feeding exclusively on earthworms in Douglas County.
Other snakes and moles are the main predators on this small creature in Kansas (Collins, 1993).

Remarks
First reported from Kansas by Cope (1875) based on a specimen from "Kansas" in the United States National Museum. The earliest existing specimen (KU 1585) from Kansas was collected in Anderson County in August of 1909 (no other associated data).
Burt and Hoyle (1935) reported on a Western Wormsnake collected under stones on a grassy hillside near a spring not far from Spring River and about 3 miles north of Baxter Springs, Cherokee County, Kansas, by Edward H. Taylor and Hobart M. Smith on 4 April 1931.

References

D'Orbigny, Charles. D. (Editor). 1843. Dictionnaire universel d'histoire naturelle, Volume 3: Caama-Clavigère. Renard, Martinet, and Company, Paris, France. 298-744 pp.

Cope, Edward D. 1875. Check-list of North American Batrachia and Reptilia; with a systematic list of the higher groups, and an essay on geographical distribution. Based on the specimens contained in the U.S. National Museum. Bulletin of the United States National Museum 1:1-104.

Jordan, David Starr. 1876. Manual of the Vertebrates of the Northern United States: Including the District East of the Mississippi River, and North of North Carolina and Tennessee, Exclusive of Marine Species. Jansen, McClurg, and Company, Chicago, Illinois.. 342 pp.

Mozley, Annie E. 1877. List of Kansas snakes in the museum of the Kansas State University. Transactions of the Kansas Academy of Science 6:34-35.

Jordan, David S. 1878. Manual of the Vertebrates of the Northern United States: Including the District East of the Mississippi River, and North of North Carolina and Tennessee, Exclusive of Marine Species. Second Edition, Revised, and Enlarged. Jansen, McClurg & Company, Chicago.

Cragin, Francis W. 1880. A preliminary catalogue of Kansas reptiles and batrachians. Transactions of the Kansas Academy of Science 7:112-123.

Yarrow, Henry C. 1882. Check list of North American Reptilia and Batrachia with catalogue of specimens in U. S. National Museum. Bulletin of the United States National Museum (24):1-249.

Garman, Samuel. 1883. The reptiles and batrachians of North America. . Kentucky Geological Survey. Yeoman Press., Frankfort, Kentucky. 185 pp.

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Brown, Arthur Erwin. 1901. A review of the genera and species of American snakes, north of Mexico. Proceedings of the Academy of Natural Sciences of Philadelphia 53(1):10-110.

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Ditmars, Raymond L. 1907. The Reptile Book; A comprehensive, Popularised Work on the Structure and Habits of the Turtles, Tortoises, Crocodilians, Lizards and Snakes which Inhabit the United States and Northern Mexico. Doubleday, Pae, and Company, New York. 472 pp.

Hurter, Julius. 1911. Herpetology of Missouri. Transactions of the Academy of Science of St. Louis 20(5):59-274.

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Linsdale, Jean M. 1925. Land Vertebrates of a Limited Area in Eastern Kansas. Thesis. University of Kansas, Lawrence, Kansas. 312 pp.

Burt, Charles E. 1927. An annotated list of the amphibians and reptiles of Riley County, Kansas. Occasional Papers of the Museum of Zoology, University of Michigan (189):12.

Linsdale, Jean M. 1927. Amphibians and reptiles of Doniphan County, Kansas. Copeia 1927(164):75-81.

Gloyd, Howard K. 1928. The amphibians and reptiles of Franklin County, Kansas. Transactions of the Kansas Academy of Science 31:115-141.

Taylor, Edward H. 1929. A revised checklist of the snakes of Kansas. University of Kansas Science Bulletin 19(5):53-62.

Gloyd, Howard K. 1932. The herpetological fauna of the Pigeon Lake Region, Miami County, Kansas. Papers of the Michigan Academy of Science, Arts and Letters 15:389-408.

Stejneger, Leonhard H. and Thomas Barbour. 1933. A Checklist of North American Amphibians and Reptiles. 3rd Edition. Harvard University Press, Cambridge, Massachusetts.

Burt, Charles E. 1933. Some distributional and ecological records of Kansas reptiles. Transactions of the Academy of Science of St. Louis 26:186-208.

Burt, Charles E. and W. L. Hoyle. 1934. Additional records of the reptiles of the central prairie region of the United States. Transactions of the Kansas Academy of Science 37:193-216.

Hurd, Myron Alec. 1936. The reptiles of Cherokee County, Kansas. Thesis. Pittsburg State University, Pittsburg, Kansas. 103 pp.

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Smith, Hobart M. 1950. Handbook of Amphibians and Reptiles of Kansas. University of Kansas, Museum of Natural History, Miscellaneous Publication (2):336.

Brumwell, Malcolm J. 1951. An ecological survey of the Fort Leavenworth Military Reservation. The American Midland Naturalist 45(1):187-231.

Schmidt, Karl P. 1953. A Check List of North American Amphibians and Reptiles. 6th Edition. University of Chicago Press, Chicago, Illinois. 280 pp.

Smith, Hobart M. 1956. Handbook of Amphibians and Reptiles of Kansas. Second edition. University of Kansas Museum of Natural History Miscellaneous Publication (9):1-356.

Loomis, Richard B. 1956. The chigger mites of Kansas (Acarina, Trombiculidae). University of Kansas Science Bulletin 37:1195-1443.

Clarke, Robert F. 1958. An ecological study of the reptiles and amphibians in Osage County, Kansas. Emporia State Research Studies 7(1):1-52.

Choate, Jerry R. 1967. Wildlife in the Wakarusa Watershed of Northeastern Kansas. Kansas Biological Survey, Lawrence. 46 pp.

Clark, Donald R., Jr. 1967. Experiments into selection of soil type, soil moisture level, and temperature by five species of small snakes. Transactions of the Kansas Academy of Science 70:490-496.

Gier, Herschel T. 1967. Vertebrates of the Flint Hills. Transactions of the Kansas Academy of Science 70(1):51-59.

Fitch, Henry S. 1970. Reproductive cycles in lizards and snakes. University of Kansas Museum of Natural History Miscellaneous Publication (52):1-247.

Clark, Donald R., Jr. 1970. Ecological study of the worm snake Carphophis vermis (Kennicott). University of Kansas Publications Museum of Natural History 19(2):213-219.

Fitch, Henry S. and Robert R. Fleet. 1970. Natural history of the milk snake (Lampropeltis triangulum) in northeastern Kansas. Herpetologica 26(4):387-396.

Aldridge, Robert D., and D. E. Metter. 1973. The reproductive cycle of the western worm snake, Carphophis vermis, in Missouri. Copeia 1973(3):472-477.

Henderson, Robert W. 1974. Resource partitioning among snakes of the University of Kansas Natural History Reservation: A preliminary analysis. Milwaukee Public Museum Contributions in Biology and Geology (1):1-11.

Collins, Joseph T. 1974. Amphibians and Reptiles in Kansas. University of Kansas Museum of Natural History Public Education Series (1):283 pp.

Pisani, George R. 1974. Herpetology in the KU Division of Biological Sciences. Kansas Herpetological Society Newsletter (4):3-4.

Karns, Daryl, Ray E. Ashton, Jr., and Thomas Swearingen. 1974. Illustrated Guide to Amphibians and Reptiles in Kansas: An Identification Manual. University of Kansas Publications Museum of Natural History Public Education Series(2):viii + 18.

Rundquist, Eric M. 1976. Field checklist (of) amphibians and reptiles of Kansas. Kansas Herpetological Society, Lawrence.

Caldwell, Janalee P. and Gregory. Glass. 1976. Vertebrates of the Woodson County State Fishing Lake and Game Management Area. Pages 62-76 in Preliminary inventory of the biota of Woodson County State Fishing Lake and Game Management Area. Report No. 5. State Biological Survey of Kansas, Lawrence.

Perry, Janice. 1977. KHS members achieve goal: Get Cottonmouth. Kansas Herpetological Society Newsletter (21):3-4.

Perry, Janice. 1978. KHS successful at Miami County State Lake. Kansas Herpetological Society Newsletter (27):5.

Collins, Joseph T. and Janalee P. Caldwell. 1978. New records of fishes, amphibians, and reptiles in Kansas for 1977. Technical Publication of the State Biological Survey of Kansas 6:70-88.

Dloogatch, M. 1978. Eggs and hatchlings of the Worm Snake, Carphophis vermis (Kennicott). Bulletin of Chicago Herpetological Society 13:99-100.

Collins, Joseph T. 1979. New records of fishes, amphibians, and reptiles in Kansas for 1978. Technical Publication of the State Biological Survey of Kansas 8:56-66.

Spencer, Dwight. 1980. Spencer, D. 1980. Ross Natural History Reservation: the first twenty years, 1959 to 1979. Emporia State University, Emporia, Kansas.. 64 pp.

Collins, Joseph T. 1982. Report to the Kansas Fish and Game Commission on the status of three amphibians in southeastern Kansas. Kansas Fish and Game Commission, Pratt. 57 pp.

Fitch, Henry S. 1982. Resources of a snake community in prairie-woodland habitat of northeastern Kansas. Pages 83-97 in Herpetological communities: A symposium of the Society for the Study of Amphibians and Reptiles and the Herpetologists League, August 1977.  Wildlife Research Reports 12. 239 pp. U. S. Fish and Wildlife Service, Washington, D. C.

Collins, Joseph T. 1982. Amphibians and Reptiles in Kansas. 2nd edition. University of Kansas Museum of Natural History Public Education Series (8).

Collins, Joseph T. 1983. New records of fishes, amphibians, and reptiles in Kansas for 1982 . Technical Publication of the State Biological Survey of Kansas 13:9-21.

Brown, Kenneth L. 1984. Pomona: A plains village variant in eastern Kansas and western Missouri. Dissertation. University of Kansas, Lawrence, Kansas. 519 pp.

Heinrich, Mark L. 1984. Herpetofauna of the Konza Prairie Research Natural Area in the Flint Hills region of Kansas with respect to habitat selection. Thesis. Kansas State University, Manhattan, Kansas. 57 pp.

Secor, Stephen M. and Charles C. Carpenter. 1984. Distribution maps of Oklahoma reptiles. Oklahoma Herpetological Society Special Publication (3):1-57.

Fitch, Henry S. 1985. Variation in clutch and litter size in New World reptiles. University of Kansas Museum of Natural History Miscellaneous Publication (76):1-76.

Lynch, John D. 1985. Annotated checklist of the amphibians and reptiles of Nebraska. Transactions of the Nebraska Academy of Sciences 13:33-57.

Heinrich, Mark L. and D. W. Kaufman. 1985. Herpetofauna of the Konza Prairie Research Natural Area, Kansas. Prairie Naturalist 17(2):101-112.

Collins, Joseph T. 1986. New records of amphibians and reptiles in Kansas for 1985. Kansas Herpetological Society Newsletter (63):4.

Collins, Joseph T. 1986. New records of amphibians and reptiles in Kansas for 1986. Kansas Herpetological Society Newsletter (66):9-16.

Coleman, Keith. 1987. Annual KHS Field Trip held at Atchison State Lake. Kansas Herpetological Society Newsletter (68):5-6.

Collins, Joseph T. 1989. New records of amphibians and reptiles in Kansas for 1989. Kansas Herpetological Society Newsletter (78):16-21.

Collins, Joseph T. 1990. Results of second Kansas herp count held during April-May 1990. Kansas Herpetological Society Newsletter (81):10-12.

Collins, Joseph T. 1990. Maximum size records for Kansas amphibians and reptiles. Kansas Herpetological Society Newsletter (81):13-17.

Joy, Jack. 1990. An additional note on Howard K. Gloyd. Bulletin of the Chicago Herpetological Society 25(10):180.

Conant, Roger and Joseph T. Collins. 1991. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed. Houghton Mifflin Company, Boston, Massachusetts.

Fitch, Henry S. 1991. Reptiles and amphibians of the Kansas ecological reserves. Pages 71-74 in Ecology and Hydrology of Kansas Ecological Reserves and the Baker Wetlands. Multidisciplinary Guidebook 4. Kansas Academy of Science, Lawrence, Kansas.

Collins, Joseph T. 1991. Results of third Kansas herp count held during April-May 1991. Kansas Herpetological Society Newsletter (85):9-13.

Collins, Joseph T. 1992. Results of the fourth Kansas herp count held during April-May 1992. Kansas Herpetological Society Newsletter (89):10-.

Collins, Joseph T. and Suzanne L. Collins. 1993. Amphibians and Reptiles in Kansas. Third Edition. University Press of Kansas, Lawrence, Lawrence. 397 pp.

Collins, Joseph T. and Rundquist, Eric M. 1993. Results of the fifth Kansas herp count held during April-June 1993 . Kansas Herpetological Society Newsletter (94):7-11.

Fitch, Henry S. 1993. Relative abundance of snakes in Kansas. Transactions of the Kansas Academy of Science 96(3/4):213-224.

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Rundquist, Eric M. 1994. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14.

Riedle, J. Daren. 1994. A survey of reptiles and amphibians at Montgomery County State Fishing Lake. Kansas Herpetological Society Newsletter (98):11-13.

Dloogatch, Michael A. (Editor). 1994. Herpetology 1994. Bulletin of the Chicago Herpetological Society 29(12):282-283.

Collins, Joseph T. 1995. New records of amphibians and reptiles in Kansas for 1994. Kansas Herpetological Society Newsletter (100):24-47.

Rundquist, Eric M. 1995. Results of the seventh annual KHS herp counts held 1 April-31 May 1995. Kansas Herpetological Society Newsletter (101):11-17.

Rundquist, Eric M. 1995. Additional KHS herp counts for 1995. Kansas Herpetological Society Newsletter (102):11-.

Rundquist, Eric M. 1996. Results of the eighth annual KHS herp counts Held 1 April-31 May 1996. Kansas Herpetological Society Newsletter (104):6-17.

Rakestraw, J. 1996. Spring herp counts: A Kansas tradition. Reptile & Amphibian Magazine (March-April):75-80.

Collins, Joseph T. 1997. A report on the KHS fall field trip to the Marais des Cygnes wildlife refuges. Kansas Herpetological Society Newsletter (110):2-3.

Conant, Roger and Joseph T. Collins. 1998. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed, expanded. Houghton Mifflin Company, Boston, Massachusetts.

Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. 1998. A Key to Amphibians & Reptiles of the Continental United States and Canada. Univ Press of Kansas, Lawrence, Kansas. 131 pp.

Gamble, Jerre. 1998. Marais des Cygnes National Wildlife Refuge Comprehensive Conservation Plan. U.S. Fish and Wildlife Service, Hartford, Kansas. 91 pp.

Rundquist, Eric M. 1998. Results of the tenth annual KHS herp counts for 1998, held 1 April-31 May. Kansas Herpetological Society Newsletter (112):11-18.

Rundquist, Eric M. 1999. Kansas Herpetological Society herp counts: A 10 year summary and evaluation. Kansas Herpetological Society Newsletter (115):42962.

Taggart, Travis W. 1999. Cherokee County fall 1999 herp count. Kansas Herpetological Society Newsletter (117):6.

Rundquist, Eric M. 2000. Results of the eleventh and twelfth annual KHS herpetofaunal counts for 1999-2000, held 1 April-31 May. Kansas Herpetological Society Newsletter (122):11-16.

Taggart, Travis W. 2001. The KHS 2001 spring field trip: A rainy rendezvous. Kansas Herpetological Society Newsletter (124):12-14.

Rundquist, Eric M. 2001. Results of the thirteenth annual KHS herp counts for 2001, held 1 April-30 June. Kansas Herpetological Society Newsletter (125):13-16.

Kingsbury, Bruce and Joanna Gibson. 2002. Habitat Management Guidelines for Amphibians and Reptiles of the Midwest. Publication of Partners in Amphibian and Reptile Conservation, Address not given. 152 pp.

Fogell, Daniel D. 2002. Occurrence and relative abundance of amphibians and reptiles at Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. Interim Report. National Park Service, Washington, D.C.. 6 pp.

Riedle, J. Daren and A. Hynek. 2002. Amphibian and reptile inventory of the Kansas Army Ammunition Plant, Labette County, Kansas. Journal of Kansas Herpetology (2):18-20.

Freeman, Craig C. 2003. A natural areas inventory of the Ft. Leavenworth Military Reservation, Leavenworth County, Kansas. II. Open-file Report No. 117. Kansas Biological Survey, Lawrence, Kansas. 199 pp.

Fogell, Daniel D. 2003. A herpetofaunal inventory of Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. National Park Service, Washington, D.C.. 59 pp.

Ernst, Carl H. and E. M. Ernst. 2003. Snakes of the United States and Canada. Smithsonian Institution Press, Washington, D. C. 668 pp.

Lokke, John L. 2003. Geographic distribution: Carphophis vermis. Journal of Kansas Herpetology (5):10.

Suleiman, G. 2003. Fort Riley herpetofaunal count. Journal of Kansas Herpetology (5):11-12.

Fitch, Henry S. 2003. Reproduction in snakes of the Fitch Natural History Reservation in northeastern Kansas. Journal of Kansas Herpetology (6):21-24.

Taggart, Travis W. 2003. Results of the 2003 KHS spring field trip to Wilson County. Journal of Kansas Herpetology (6):2-5.

Ernst, Carl H., J. M. Orr, and T. R. Creque. 2003. Carphophis. Catalogue of American Amphibians and Reptiles (773):1-2.

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Collins, Joseph T. 2004. New records of amphibians, turtles, and reptiles in Kansas for 2003. Journal of Kansas Herpetology (9):8-11.

Daniel, James K. 2004. Cherokee County herp count. Journal of Kansas Herpetology (11):10.

Collins, Joseph T. 2004. Marais des Cygnes herp count. Journal of Kansas Herpetology (11):11.

Willson, John D. and Michael E. Dorcas. 2004. Aspects of the ecology of small fossorial snakes in the western piedmont of North Carolina. Southeastern Naturalist 3(1):1-12.

Taggart, Travis W. 2005. Results of the KHS 2005 fall field trip [to Crawford County]. Journal of Kansas Herpetology (16):19-21.

Taggart, Travis W. 2006. Distribution and status of Kansas herpetofauna in need of information. State Wildlife Grant T7. Kansas Department of Wildlife and Parks, Pratt. vii + 106 pp.

Wilgers, Dustin J. and Eva A. Horne. 2006. Effects of different burn regimes on tallgrass prairie herpetofaunal species diversity and community composition in the Flint Hills, Kansas. Journal of Herpetology 40:73-84.

Pisani, George R. 2007. New Kansas maximum sizes for Virginia valeriae and Carphophis vermis. Journal of Kansas Herpetology (22):11.

Orr, J. M. 2007. Microhabitat use by the Eastern Worm Snake, Carphophis amoenus. Herpetological Bulletin 97:29-35.

Taggart, Travis W. 2008. KHS 2008 spring field trip. Journal of Kansas Herpetology (25):2-3.

Murrow, Daniel G. 2009. KHS 2009 spring field trip. Journal of Kansas Herpetology (29):42769.

Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. 2010. Amphibians, Reptiles, and Turtles of Kansas. Eagle Mountain Publishing., Provo, Utah. 400 pp.

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Gray, Brian S. 2010. Distribution of native and exotic earthworms in the eastern United States: Implications for the ecology of vermivorous snakes. Bulletin of the Chicago Herpetological Society 45(5):73-86.

McMartin, David C. 2011. U. S. Army 2011 Fort Leavenworth Herpetofaunal Survey: 23 April - 09 May 2011. Privately printed, Leavenworth, Kansas. 33 pp.

Taggart, Travis W. 2011. Kansas Herpetological Society 2011 spring field trip to be held in Chautauqua County. Journal of Kansas Herpetology (37):5-7.

McMartin, D. Chris. 2011. Herp Count: Fort Leavenworth Herpetofaunal Survey for 2011. Journal of Kansas Herpetology (39):8-9.

Rohweder, Megan R. 2012. Spatial conservation prioritization of Kansas for terrestrial vertebrates. Thesis. Fort Hays State University, Hays, Kansas. 151 pp.

Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. 2012. Key to the Herpetofauna of the Continental United States and Canada: Second Edition, Revised and Updated. Univ Press of Kansas, Lawrence, Kansas. 152 pp.

Hamilton, Bryan Tyler, Rachel Hart, and Jack W. Sites Jr. 2012. Feeding ecology of the Milksnake (Lampropeltis triangulum, Colubridae) in the western United States. Journal of Herpetology 46(4):515-522.

Sinclair, Tom. 2013. A four-day spring snake count across northern Kansas. Collinsorum 2(1/2):9.

Taggart, Travis W. 2013. KHS 2012 Spring Field Trip to Bourbon County State Lake. Collinsorum 2(3/4):3.

Taggart, Travis W. 2013. KHS 2013 Spring Field Trip to Schermerhorn Park, Cherokee County. Collinsorum 2(3/4):4.

McMartin, D. Chris. 2014. Fort Leavenworth Heretofaunal Survey for 2013. Collinsorum 3(1):10.

Pittman, Galen L., Henry S. Fitch, and W. Dean Kettle. 2016. Vertebrate animals on the Fitch Natural History Reservation (1948-2002). Kansas Biological Survey Report Number 188, Lawrence. 48 pp.

Powell, Robert, Roger Conant, and Joseph T. Collins. 2016. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. Houghton Mifflin Harcourt, Boston. 494 pp.

Crother, Brian I. (editor). 2017. Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, with Comments Regarding Confidence in Our Understanding. Eighth edition. Herpetological Circulars (43):1-102.

Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. 2019. Key to the Herpetofauna of the Continental United States and Canada. Third Edition. Univ Press of Kansas, Lawrence, Kansas. 192 pp.

Baggett, Brianna A. 2019. Phylogeography and population demography of the Eastern Worm Snake Carphophis amoenus. Thesis. University of West Georgia, Carrollton, Georgia. 47 pp.

Daniel, Richard E. and Brian S. Edmond. 2020. Atlas of Missouri Amphibians and Reptiles for 2019. Privately printed, Columbia, Missouri. 86 pp.

Riedle, J. Daren. 2020. Revisiting Kansas Herpetological Society field trip and Herp Count data: Distributional patterns and trend data of Kansas amphibians and reptiles. Collinsorum 9(1):7-16.

Hullinger, Allison, Zackary Cordes, Daren Riedle, and William Stark. 2020. Habitat assessment of the Broad-headed Skink (Plestiodon laticeps) and the associated squamate community in eastern Kansas. Transactions of the Kansas Academy of Science 123(1-2):137-150.

Taggart, Travis W and Sarah L Taggart. 2021. Herp Count: Cherokee County: KHS-2020-02. Collinsorum 9(3):11-12.

Taggart, Travis W and Sarah L Taggart. 2021. Herp Count: Cherokee County: KHS-2020-03. Collinsorum 9(3):12.

Riedle, J. Daren, Tamera D. Riedle, Zachary Riedle, and Greya Riedle. 2021. Herp Count: Montgomery County: KHS-2020-23. Collinsorum 9(3):14-15.

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THE CENTER FOR NORTH AMERICAN HERPETOLOGY — Accessed: Tuesday 09 June 2026 13:13 CT