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Eastern Copperhead

Agkistrodon contortrix (Linnaeus, 1766)

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Overview
DANGEROUSLY VENOMOUS (a threat to life or limb): The Eastern Copperhead is the only distinctly banded snake in Kansas with no rattle on its tail and a small pit on each side of its head between and slightly below the eye and nostril. This snake normally has a pattern of 7- 20 light-edged crossbands on its body; the bands are narrow on the back and wide on the sides.
Eastern Copperheads vary in color from gray to light brown with dark gray or brown crossbands. The head may be gray, brown, or reddish. The belly is white, with large dark gray, brown, or black blotches along its edges that extend a short distance up onto the sides of the body.
Young specimens have yellowish tails. Adult male Eastern Copperheads grow much longer than females. Fitch (1959) discovered several male Eastern Copperheads in Douglas County with no crossband pattern; the snakes were uniform, dull brown. Young look like miniature adults, but with yellowish or greenish tails.
The two species hybridize across much of Kansas.
Adults normally grow to 56.0-91.5 cm (22-36 inches) in total length. The largest specimen from Kansas is a male (KU 196643) from Jefferson County with a total length of 102.0 cm (40 inches), collected by Henry S. Fitch on 13 July 1984. The maximum length throughout the range is 134.6 cm (53 inches) (Powell et al., 2016). The maximum weight for a Kansas specimen is about 400 grams (14 ounces).

Distribution
The two forms are broadly overlapping. Agkistrodon contortrix is found generally north and east of the Neosho River basin, where it is scarce and occurs in scattered populations. It enters the Flint Hills along rocky riparian corridors where there has been genetic admixture at some point in the past with the Broad-banded Copperhead.
Hybrids occur primarily between the Verdigris River and Walnut River drainages in Kansas.
Additionally, genetic hybrids were identified among the westernmost populations of the Eastern Copperhead (A. contortrix) along the Kansas and Neosho River drainages (Burbrink and Guiher, 2014).
Two 'pure' Eastern Copperheads were detected in Elk County, and these outlier samples should be reexamined.
Branson (1904) reported examining a specimen from Rook County; the locality is undoubtedly in error.

Reproduction
Eastern Copperheads are found in open rocky woodland, woodland edge, and meadows with clumps of brush adjacent to woodland. Their pattern and color blend perfectly with forest leaf litter. This species is active from April to early November and prefers an optimal temperature near 80°F. During the cool spring and fall months, this snake is active only during the day and is normally found coiled on rocky hillsides or forest floors, awaiting the approach of prey.
In the summer, Eastern Copperheads become nocturnal and prowl for food, aided by their night vision and the two sensory pits between the eyes and nostrils. These pits detect the body heat of warm-blooded prey such as mice. Eastern Copperheads have home ranges, but males tend to wander in larger areas than females. Home ranges vary from eight to 25 acres. Because of their shy, retiring disposition and camouflage pattern, these snakes easily exist in reasonably large numbers near areas of heavy human population. Estimates indicate a population density of three or four Eastern Copperheads per acre in northeastern Kansas (Fitch, 1982). The few people bitten by these snakes each year are usually victims of accidents, not aggressive, dangerous animals. During the fall, this snake returns to the wooded hilltop rock outcrops with a southern exposure that it used in previous years and retires deep beneath the ground to avoid the cold temperatures of winter.
This snake breeds from April to August, with a peak of activity in April and May, just after spring emergence from winter inactivity. Adult male Eastern Copperheads are among those snakes that infrequently engage in combat dances. A typical combat dance involves two males facing each other with their heads and the front of their bodies raised off the ground. They come together so their bellies are firmly adpressed. Then they slowly. intertwine their necks with writhing movements, until suddenly one male hurls the other to the ground with a quick body contraction. The combat dance evidently establishes dominance between two males. Many female Eastern Copperheads bear young only every other year. Courtship and mating have not been adequately observed and are presumably nocturnal. The young are born venomous and number from one to fourteen per litter (Fitch, 1985), with an average of about five. Females give birth in August, September, and October. Species of Agkistrodon are viviparous and develop functional chorioallantoic and yolk-sac (omphalallantoic) placentae. Embryos are predominantly lecithotrophic, but placental exchange supports the exchange of respiratory gases, water, and inorganic ions. Experimental isotope work (e.g., in A. contortrix) demonstrates maternal transfer of dietary amino acids during gestation, consistent with incipient placentotrophy, although yolk remains the primary source of embryonic dry mass.
Eastern Copperheads feed on insects (particularly cicadas), frogs, toads, lizards, small birds, and other snakes. They are particularly fond of rodents. Young Eastern Copperheads may use their yellowish tails as lures to attract prey such as small frogs and toads.
Predators of Eastern Copperheads include large birds, mammals, and larger snakes, but people are its chief enemy.

Remarks
First reported in Kansas by Hallowell (1857) based on specimens collected by William A. Hammond near Fort Riley, Riley County. The earliest extant specimen (MCZ R5844) was collected at "Ft Riley, Kan." and received from the United States National Museum in 1879.
Specimens are needed along the western periphery of its range in Kansas to better define its limits there. Additional specimens are also desirable from the Drift Hills in the northeast and throughout the Neosho River drainage basin.
Burt (1935) reported a specimen collected 5 miles east of Rock, Cowley County, Kansas, on 16 April 1934.
Gloyd (1969) described Agkistrodon contortrix phaeogaster. United States National Museum no. 165955, adult male, received from Dr. Henry S. Fitch, July 1960; locality 10 miles south of McLouth, Jefferson County, Kansas; collector not stated. He went on to define its distribution as "A. c. phaeogaster occupies the major portion of the Marais des Cygnes-Osage River drainage system in eastern Kansas and central Missouri. It intergrades with A. c. laticinctus in southern Kansas and northern Oklahoma... ".
Burbrink and Guiher (2014) recognized two species A. contortrix (eastern-most populations) and A. laticinctus (western populations) in the United States. The two species mix over eastern Texas and eastern Oklahoma. And, based on relatively extensive sampling in Kansas, they discovered that the entire range in Kansas was an admixture of the two species. Because the two species cannot be differentiated in Kansas without genetic analysis, the complex has been grouped into a single account.
Vial et al. (1977), using data from Eastern Copperheads studied in northeastern Kansas, estimated a natural longevity of up to 15 years for this snake. Based on a captive specimen, Snider and Bowler (1992) reported a maximum longevity of 29 years, 10 months, and 6 days for this species.

References

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Smith, Hobart M. 1950. Handbook of Amphibians and Reptiles of Kansas. University of Kansas, Museum of Natural History, Miscellaneous Publication (2):336.

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Smith, Hobart M. 1956. Handbook of Amphibians and Reptiles of Kansas. Second edition. University of Kansas Museum of Natural History Miscellaneous Publication (9):1-356.

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McDuffie, George T. 1960. Studies on the ecology and life history of the Copperhead, Agkistrodon contortrix mokeson (Daudin), in Ohio. Dissertation. University of Cincinnati, Cincinnati, Ohio. 82 pp.

Fitch, Henry S. 1960. Autecology of the copperhead. University of Kansas Publications Museum of Natural History 13(4):85-288.

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Caldwell, Janalee P. and Gregory. Glass. 1976. Vertebrates of the Woodson County State Fishing Lake and Game Management Area. Pages 62-76 in Preliminary inventory of the biota of Woodson County State Fishing Lake and Game Management Area. Report No. 5. State Biological Survey of Kansas, Lawrence.

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Perry, Janice. 1977. KHS members achieve goal: Get Cottonmouth. Kansas Herpetological Society Newsletter (21):3-4.

Curl, Richard L. 1978. Final Environmental Statement: Milford Lake Kansas operation and maintenance. US Army Corps of Engineers, Kansas City District. 158 pp.

Fitch, Henry S. and E. Raymond Hall. 1978. A 20year record of succession on reseeded fields of tallgrass prairie on the Rockefeller Experimental Tract. University of Kansas Museum of Natural History Special Publication (4):1-15.

Capron, Marty B. 1978. Notes on the possible occurrence of Broadbanded Copperheads in Kansas. Kansas Herpetological Society Newsletter (26):4-6.

Perry, Janice. 1978. KHS successful at Miami County State Lake. Kansas Herpetological Society Newsletter (27):5.

Collins, Joseph T. 1978. The Osage Copperhead. University of Kansas Museum of Natural History Association Newsleter (7(2)):71-74.

Collins, Joseph T. and Janalee P. Caldwell. 1978. New records of fishes, amphibians, and reptiles in Kansas for 1977. Technical Publication of the State Biological Survey of Kansas 6:70-88.

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Smith, Donald D. 1980. The Copperhead. AALAS Kansas City Branch News 3(2):8-10.

Lokke, John L. 1981. Featured herp: Agkistrodon contortrix phaeogaster. Nebraska Herpetological Society Newsletter 3(1):4-5.

Fitch, Henry S. 1982. Resources of a snake community in prairie-woodland habitat of northeastern Kansas. Pages 83-97 in Herpetological communities: A symposium of the Society for the Study of Amphibians and Reptiles and the Herpetologists League, August 1977.  Wildlife Research Reports 12. 239 pp. U. S. Fish and Wildlife Service, Washington, D. C.

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Miller, Larry L. 1983. Bourbon County field trip well attended and successful. Kansas Herpetological Society Newsletter (54):6-7.

Collins, Joseph T. 1983. New records of fishes, amphibians, and reptiles in Kansas for 1982 . Technical Publication of the State Biological Survey of Kansas 13:9-21.

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Brown, Kenneth L. 1984. Pomona: A plains village variant in eastern Kansas and western Missouri. Dissertation. University of Kansas, Lawrence, Kansas. 519 pp.

Heinrich, Mark L. 1984. Herpetofauna of the Konza Prairie Research Natural Area in the Flint Hills region of Kansas with respect to habitat selection. Thesis. Kansas State University, Manhattan, Kansas. 57 pp.

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Collins, Joseph T. 1984. New records of fishes, amphibians and reptiles in Kansas for 1984. Kansas Herpetological Society Newsletter (58):14-20.

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Fitch, Henry S. and Joseph T. Collins. 1985. Intergradation of the Osage and Broad-banded Copperheads in Kansas. Transactions of the Kansas Academy of Science 88(3-4):135-137.

Collins, Joseph T. 1986. New records of amphibians and reptiles in Kansas for 1986. Kansas Herpetological Society Newsletter (66):9-16.

Coleman, Keith. 1987. Annual KHS Field Trip held at Atchison State Lake. Kansas Herpetological Society Newsletter (68):5-6.

Collins, Joseph T. 1989. New records of amphibians and reptiles in Kansas for 1989. Kansas Herpetological Society Newsletter (78):16-21.

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Collins, Joseph T. 1990. Results of second Kansas herp count held during April-May 1990. Kansas Herpetological Society Newsletter (81):10-12.

Collins, Joseph T. 1990. Maximum size records for Kansas amphibians and reptiles. Kansas Herpetological Society Newsletter (81):13-17.

Joy, Jack. 1990. An additional note on Howard K. Gloyd. Bulletin of the Chicago Herpetological Society 25(10):180.

Conant, Roger and Joseph T. Collins. 1991. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed. Houghton Mifflin Company, Boston, Massachusetts.

Fitch, Henry S. 1991. Reptiles and amphibians of the Kansas ecological reserves. Pages 71-74 in Ecology and Hydrology of Kansas Ecological Reserves and the Baker Wetlands. Multidisciplinary Guidebook 4. Kansas Academy of Science, Lawrence, Kansas.

Collins, Joseph T. 1991. New records of amphibians and reptiles in Kansas for 1990. Kansas Herpetological Society Newsletter (83):7-13.

Collins, Joseph T. 1991. Results of third Kansas herp count held during April-May 1991. Kansas Herpetological Society Newsletter (85):9-13.

Collins, Joseph T. 1991. Viewpoint: A new taxonomic arrangement for some North American amphibins and reptiles. Herpetological Review 22(2):42-43.

Ernst, Carl H. 1992. Venomous Reptiles of North America. Smithsonian Institution Press, Washington, D. C.

Knight, A., L. D. Densmore III, and E. D. Rael. 1992. Molecular systematics of the Agkistrodon complex. Pages 49-69 in Biology of the Pitvipers. Selva, Tyler, Texas.

Collins, Joseph T. 1992. Results of the fourth Kansas herp count held during April-May 1992. Kansas Herpetological Society Newsletter (89):10-.

Collins, Joseph T. and Suzanne L. Collins. 1993. Amphibians and Reptiles in Kansas. Third Edition. University Press of Kansas, Lawrence, Lawrence. 397 pp.

Collins, Joseph T. and Rundquist, Eric M. 1993. Results of the fifth Kansas herp count held during April-June 1993 . Kansas Herpetological Society Newsletter (94):7-11.

Fitch, Henry S. 1993. Relative abundance of snakes in Kansas. Transactions of the Kansas Academy of Science 96(3/4):213-224.

Fitzgerald, Eve C. and Charles Nilon. 1994. Classification of habitats for endangered and threatened species in Wyandotte County, Kansas. Kansas Department of Wildlife and Parks, Pratt, Kansas. 98 pp.

Rundquist, Eric M. 1994. 1994 Field Trip. Kansas Herpetological Society Newsletter (95):3-4.

Rundquist, Eric M. 1994. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14.

Riedle, J. Daren. 1994. A survey of reptiles and amphibians at Montgomery County State Fishing Lake. Kansas Herpetological Society Newsletter (98):11-13.

Dloogatch, Michael A. (Editor). 1994. Herpetology 1994. Bulletin of the Chicago Herpetological Society 29(12):282-283.

Holman, J. Alan. 1995. Pleistocene Amphibians and Reptiles. Oxford University Press, New York. 243 pp.

Rundquist, Eric M. 1995. Results of the seventh annual KHS herp counts held 1 April-31 May 1995. Kansas Herpetological Society Newsletter (101):11-17.

Rundquist, Eric M. 1995. Additional KHS herp counts for 1995. Kansas Herpetological Society Newsletter (102):11-.

Parmley, Dennis and J. Alan Holman. 1995. Hemphillian (Late Miocene) snakes from Nebraska, with comments on Arikareean through Blancan Snakes of midcontinental North America. Journal of Vertebrate Paleontology 15(1):79-95.

Rundquist, Eric M. 1996. Results of the eighth annual KHS herp counts Held 1 April-31 May 1996. Kansas Herpetological Society Newsletter (104):6-17.

Rakestraw, J. 1996. Spring herp counts: A Kansas tradition. Reptile & Amphibian Magazine (March-April):75-80.

Rundquist, Eric M. 1997. Results of the ninth annual KHS herp counts held 1 April-31 May 1997. Kansas Herpetological Society Newsletter (108):12-17.

Collins, Joseph T. 1997. A report on the KHS fall field trip to the Marais des Cygnes wildlife refuges. Kansas Herpetological Society Newsletter (110):2-3.

Conant, Roger and Joseph T. Collins. 1998. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed, expanded. Houghton Mifflin Company, Boston, Massachusetts.

Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. 1998. A Key to Amphibians & Reptiles of the Continental United States and Canada. Univ Press of Kansas, Lawrence, Kansas. 131 pp.

Gamble, Jerre. 1998. Marais des Cygnes National Wildlife Refuge Comprehensive Conservation Plan. U.S. Fish and Wildlife Service, Hartford, Kansas. 91 pp.

Rundquist, Eric M. 1998. Results of the tenth annual KHS herp counts for 1998, held 1 April-31 May. Kansas Herpetological Society Newsletter (112):11-18.

Schmidt, Curtis J. 1998. Herpetological observations at Cheyenne Bottoms, Barton County, Kansas. Kansas Herpetological Society Newsletter (113):15.

Collins, Joseph T. 1998. Results of the KHS silver anniversary fall field trip. Kansas Herpetological Society Newsletter (114):6-.

Rundquist, Eric M. 1999. Kansas Herpetological Society herp counts: A 10 year summary and evaluation. Kansas Herpetological Society Newsletter (115):42962.

Taggart, Travis W. 1999. Cherokee County fall 1999 herp count. Kansas Herpetological Society Newsletter (117):6.

Fitch, Henry S. 2000. Population structure and biomass of some common snakes in central North America. Scientific Papers of the Natural History Museum, University of Kansas (17):1-7.

Rundquist, Eric M. 2000. Results of the eleventh and twelfth annual KHS herpetofaunal counts for 1999-2000, held 1 April-31 May. Kansas Herpetological Society Newsletter (122):11-16.

Taggart, Travis W. 2000. Results of the KHS 2000 fall field trip. Kansas Herpetological Society Newsletter (122):6-8.

Johnson, Richard W. 2001. Spatial ecology of the Eastern Coachwhip (Masticophis flagellum flagellum) in and eastern Texas upland community. Thesis. Stephen F. Austin State University, Nacogdoches, Texas. 54 pp.

Kettle, W. Dean, R. H. Hagen, F. deNoyelles, Jr., and E. A. Martinko. 2001. The University of Kansas field station and ecological reserves: A half century of research and education. Miscellaneous Publication Number 9. Kansas Biological Survey, Lawrence. 68 pp.

Guarisco, Hank. 2001. Ode to an ophidion autumn. Kansas Herpetological Society Newsletter (123):19.

Kingsbury, Bruce and Joanna Gibson. 2002. Habitat Management Guidelines for Amphibians and Reptiles of the Midwest. Publication of Partners in Amphibian and Reptile Conservation, Address not given. 152 pp.

Fogell, Daniel D. 2002. Occurrence and relative abundance of amphibians and reptiles at Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. Interim Report. National Park Service, Washington, D.C.. 6 pp.

Fitch, Henry S. and Anna L. Clark. 2002. An exceptionally large natural assemblage of female Copperheads (Agkistrodon contortrix). Herpetological Review 33(2):94-95.

Fogell, Daniel D. 2003. A herpetofaunal inventory of Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. National Park Service, Washington, D.C.. 59 pp.

Suleiman, G. 2003. Fort Riley herpetofaunal count. Journal of Kansas Herpetology (5):11-12.

Taggart, Travis W. 2003. KHS conducts first systematic road survey. Journal of Kansas Herpetology (6):11-12.

Fitch, Henry S. 2003. Reproduction in snakes of the Fitch Natural History Reservation in northeastern Kansas. Journal of Kansas Herpetology (6):21-24.

Taggart, Travis W. 2003. Results of the 2003 KHS spring field trip to Wilson County. Journal of Kansas Herpetology (6):2-5.

Suleiman, Gibran. 2003. Fort Riley herp count. Journal of Kansas Herpetology (7):9.

Taggart, Travis W. 2003. Results of the KHS 2003 fall field trip. Journal of Kansas Herpetology (8):14-15.

Fitch, Henry S., Scott Sharp, and Kylee Sharp. 2003. Snakes of the University of Kansas biotic succession area. Journal of Kansas Herpetology (8):20-21.

Fitch, Henry S. 2004. The effect of female size on number of eggs or young in snakes. Journal of Kansas Herpetology (9):11-12.

Fitch, Henry S. 2004. Food surplus and body size in local populations of snakes. Journal of Kansas Herpetology (10):14-16.

Volkmann, Al. 2004. Cowley County herp count. Journal of Kansas Herpetology (11):10.

Fitch, Henry S. 2004. Observations on Osage Copperheads in northeastern Kansas. Journal of Kansas Herpetology (12):20.

Dloogatch, Michael A. (Editor). 2004. Herpetology 2004. Bulletin of the Chicago Herpetological Society 39(9):175-178.

Taggart, Travis W. 2006. Distribution and status of Kansas herpetofauna in need of information. State Wildlife Grant T7. Kansas Department of Wildlife and Parks, Pratt. vii + 106 pp.

Wozniak, Edward J., John Wisser, and Michael Schwartz. 2006. Venomous adversaries: A reference to snake identification, field safety, and bite-victim first aid for disaster-response personnel deploying into the hurricane-prone regions of North America. Wilderness and Environmental Medicine 17:246 266.

Maher, B. W. and Gregory Sievert. 2006. Agkistrodon contortrix. Geographic distribution. Herpetological Review 37:496.

Wilgers, Dustin J., Eva A. Horne, Brett K. Sandercock, and Allan W. Volkmann. 2006. Effects of rangeland management on community dynamics of the herpetofauna of the tallgrass prairie. 62:378-388.

Taggart, Travis W. 2008. KHS 2008 spring field trip. Journal of Kansas Herpetology (25):2-3.

Nolting, Ray. 2008. Snake hunter finds rattlesnake in Neosho. Parsons Sun 1 May 2008:1,8.

Murrow, Daniel G. 2009. KHS 2009 spring field trip. Journal of Kansas Herpetology (29):42769.

Douglas, Michael E., Marlis R. Douglas, Gordon W. Schuett, and Louis W. Porras. 2009. Climate change and evolution of the New World pitviper genus Agkistrodon (Viperidae). Journal of Biogeography 36:1164-1180.

Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. 2010. Amphibians, Reptiles, and Turtles of Kansas. Eagle Mountain Publishing., Provo, Utah. 400 pp.

Rodriguez, Paul J. 2010. Translocation of the Copperhead (Agkistrodon contortrix) in Nebraska. Thesis. University of Nebraska, Lincoln, Nebraska. 83 pp.

Murrow, Daniel G. 2010. Kansas Herpetological Society spring field trip. Journal of Kansas Herpetology (33):2-3.

Ernst, Carl H. and Evelyn M. Ernst. 2011. Venomous Reptiles of the United States, Canada, and Northern Mexico. Volume 1. Heloderma, Micruroides, Micrurus, Pelamis, Agkistrodon, Sistrurus. Johns hopkins University Press, Baltimore, Maryland.

McMartin, David C. 2011. U. S. Army 2011 Fort Leavenworth Herpetofaunal Survey: 23 April - 09 May 2011. Privately printed, Leavenworth, Kansas. 33 pp.

Taggart, Travis W. 2011. Kansas Herpetological Society 2011 spring field trip to be held in Chautauqua County. Journal of Kansas Herpetology (37):5-7.

Taggart, Travis W. 2011. Results of the KHS Spring Field Trip to Chautauqua County. Journal of Kansas Herpetology (38):2-4.

Houck, Mike. 2011. Fort Riley Herpetofaunal Survey for 2011. Journal of Kansas Herpetology (39):9.

Rohweder, Megan R. 2012. Spatial conservation prioritization of Kansas for terrestrial vertebrates. Thesis. Fort Hays State University, Hays, Kansas. 151 pp.

Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. 2012. Key to the Herpetofauna of the Continental United States and Canada: Second Edition, Revised and Updated. Univ Press of Kansas, Lawrence, Kansas. 152 pp.

Sinclair, Tom. 2013. A four-day spring snake count across northern Kansas. Collinsorum 2(1/2):9.

Taggart, Travis W. 2013. KHS 2012 Spring Field Trip to Bourbon County State Lake. Collinsorum 2(3/4):3.

Taggart, Travis W. 2013. KHS 2012 Fall Field Trip to Atchison County State Lake. Collinsorum 2(3/4):4.

Taggart, Travis W. 2013. KHS 2013 Spring Field Trip to Schermerhorn Park, Cherokee County. Collinsorum 2(3/4):4.

Taggart, Travis W. 2013. KHS 2013 Fall Field Trip to Butler County State Lake. Collinsorum 2(3/4):6.

Mardis, Dexter and Kevin Scott. 2013. 2013 Kansas Herpetofaunal Counts. Collinsorum 2(3/4):7.

McMartin, D. Chris. 2014. Fort Leavenworth Heretofaunal Survey for 2013. Collinsorum 3(1):10.

Taggart, Travis W. 2014. Results of the 2014 KHS Fall Field Trip to Woodson County. Collinsorum 3(2-4):12.

Taggart, Travis W. 2014. Recent scientific and standard English name changes effecting the Kansas herpetofauna. Collinsorum 3(2-4):9-10.

Burbrink, Frank T. and Timothy J. Guiher. 2014. Considering gene flow when using coalescent methods to delimit lineages of North American pitvipers of the genus Agkistrodon. Zoological Journal of the Linnean Society 2014:1-22.

Houck, Mike. 2015. 2015 Fort Riley Herpetofaunal Count final report. Collinsorum 4(1):10-11.

Burbrink, Frank T. and Timothy J. Guiher. 2015. Considering gene flow when using coalescent methods to delimit lineages of North American pitvipers of the genus Agkistrodon. Zoological Journal of the Linnean Society 173:505–526.

Pittman, Galen L., Henry S. Fitch, and W. Dean Kettle. 2016. Vertebrate animals on the Fitch Natural History Reservation (1948-2002). Kansas Biological Survey Report Number 188, Lawrence. 48 pp.

Powell, Robert, Roger Conant, and Joseph T. Collins. 2016. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. Houghton Mifflin Harcourt, Boston. 494 pp.

Petersen, Christopher, Robert E. Lovich, and Sarah Stallings. 2016. Herpetofauna Biodiversity On United States Army Installations. Final Report. Department of Defense Legacy Resource Management Program, Arlington, Virginia. 82 pp.

Taggart, Travis W. 2016. Results of the KHS Summer field trip to Caney River, Chautauqua County, Kansas. Collinsorum 5(2-3):4-5.

Crother, Brian I. (editor). 2017. Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, with Comments Regarding Confidence in Our Understanding. Eighth edition. Herpetological Circulars (43):1-102.

Riedle, Daren. 2018. Conservation conversations: The year of the snake. Kansas Wildlife and Parks Magazine (November/December):6.

Houck, Mike. 2018. Herp Count: Fort Riley Military Installation. Collinsorum 7(1):17.

Petersen, Christopher E., Robert E. Lovich, and Sarah Stallings. 2018. Amphibians and reptiles of United States Department of Defense installations. Herpetological Conservation and Biology 13(3):652–661.

Leao, Suelem Muniz, Eric R. Pianka, and Nicolás Pelegrin. 2018. Is there evidence for population regulation in amphibians and reptiles? Journal of Herpetology 52(1):28-33.

Pisani, George R. and J. Daren Riedle. 2018. Ontogenetic changes in tail-length and the possible relation to caudal luring in northeast Kansas Copperheads, Agkistrodon contortrix. Transactions of the Kansas Academy of Science 121(3/4):403-410.

Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. 2019. Key to the Herpetofauna of the Continental United States and Canada. Third Edition. Univ Press of Kansas, Lawrence, Kansas. 192 pp.

Daniel, Richard E. and Brian S. Edmond. 2020. Atlas of Missouri Amphibians and Reptiles for 2019. Privately printed, Columbia, Missouri. 86 pp.

Riedle, J. Daren. 2020. Revisiting Kansas Herpetological Society field trip and Herp Count data: Distributional patterns and trend data of Kansas amphibians and reptiles. Collinsorum 9(1):7-16.

Hullinger, Allison, Zackary Cordes, Daren Riedle, and William Stark. 2020. Habitat assessment of the Broad-headed Skink (Plestiodon laticeps) and the associated squamate community in eastern Kansas. Transactions of the Kansas Academy of Science 123(1-2):137-150.

Taggart, Travis W and Sarah L Taggart. 2021. Herp Count: Cherokee County: KHS-2020-02. Collinsorum 9(3):11-12.

Taggart, Travis W and Sarah L Taggart. 2021. Herp Count: Cherokee County: KHS-2020-03. Collinsorum 9(3):12.

Burbrink, Frank and Sara Ruane. 2021. Contemporary philosophy and methods for studying speciation and delimiting species. Ichthyology & Herpetology 109(3):874-894.

Holding, Matthew L., Jason L. Strickland, Rhett M. Rautsaw, Erich P. Hofmann, Andrew J. Mason, Michael P. Hogan, Gunnar S. Nystrom, Schyler A. Ellsworth, Timothy J. Colston, Miguel Borja, Gamaliel Castaneda-Gaytan, Christoph I. Grunwald, Jason M. Jones, Luciana A. Freitas-de-Sousa, Vincent Louis Viala, Mark J. Margres, Erika Hingst-Zaher, Inacio L. M. Junqueira-de-Azevedo, Ana M. Moura-da-Silvaf, Felipe G. Grazziotin, H. Lisle Gibbs, Darin R. Rokyta, and Christopher L. Parkinson. 2021. Phylogenetically diverse diets favor more complex venoms in North American pitvipers. Proceedings of the Academy of Natural Sciences of the United States of America 118(17):10.

Petersen, Christopher E. and Robert E. Lovich. 2023. Department of Defense Partners in Amphibian and Reptile Conservation: An annotated coloring book. US Navy, Norfolk, Virginia. 24 pp.

Last Updated: 02/19/2026 8:14:08 AM CT

THE CENTER FOR NORTH AMERICAN HERPETOLOGY — Accessed: Tuesday 09 June 2026 13:12 CT